The Upper Liassic Heteroptera of Mecklenburg and Saxony

Abstract

Twenty-seven species of Heteroptera from the Upper Iiassic of Dobbertin, Mecklenburg, and East Lower Saxony are redescribed and reclassified as follows. PROGONOCIMICIDAE (= Actinoscytinidae and Eocimicidae) :Progonocimex jurassicus, P.liasinus (= Eocimex liasinus); Eocercopis ancyloptera, E. similis (= Cercoprisca similis); Archicercopis falcata. ARCHEGOCIMICIDAE (= Eonabidae and Diatillidae): Archegocimex geinitzi, A.primitiva (=Eonabis primitive), A.1iadis (= Archegocoris liadis)

New fossil Progonocimicidae (Hemiptera: Coleorrhyncha: Progonocimicoidea) from the Upper Mesozoic of northeastern China, with a phylogeny of Coleorrhyncha

“…Only Progonocimicidae are known from the Triassic. A number of Progonocimicidae fossils have also been found in Europe and Asia from the Jurassic and Cretaceous periods (Handlirsch, 1939;Popov & Wootton, 1977;Popov, 1982Popov, , 1985Popov & Shcherbakov, 1991;Szwedo, 2011;Dong et al, 2012Dong et al, , 2013. Karabasiidae fossils are known from the early Jurassic to Early Cretaceous periods (Martynov, 1926;Popov, 1985Popov, , 1989Lin, 1986;Popov & Shcherbakov, 1991).…”

“…Peloridiidae, the only extant family, was first placed in Heteroptera (Breddin, 1897), but later transferred to Homoptera (Myers & China, 1929;China, 1962;Evans, 1963). Peloridiidae and Heteroptera were treated as sister groups by Schlee (1969), Handlirsch, 1939 Germany J1 a Cicadocoris abditus Popov, 1982 Kyrgyzstan J1/J2 a C. affinis Popov, 1982 Kyrgyzstan J1 a C. ambiguous Popov, 1985 Kyrgyzstan J1/J2 a C. anglicus Popov et al, 1994 UK J1 a C. arcuatus Popov, 1982 Kyrgyzstan J1 C. assimilis Dong, Yao & Ren, 2013 China J2 a C. brunneus Hong, 1983 China J2 a C. cognatus Popov, 1982 Kyrgyzstan J1/J2 a C. fidus Popov, 1982 Kyrgyzstan J1/J2 a C. frater Popov, 1982 Kyrgyzstan J1 a C. incompletus Popov, 1985 Kyrgyzstan J1 a C. intermedius Popov, 1985 Russia J1 C. kuliki Becker-Migdisova, 1958 Kyrgyzstan T3 a C. kuzbasicus Popov, 1985 Russia J1 C. lini Wang, Szwedo & Zhang, 2009 China J2 a C. liliputus Popov, 1988 Russia J1/J2 a C. minutus Popov, 1982 Kyrgyzstan J1 a C. modestus Popov, 1985 Kyrgyzstan J1/J2 a C. paulinus Popov, 1982 Kyrgyzstan J1/J2 a C. sinensis Hong, 1983 China J2 C. varians Dong, Yao & Ren, 2012 China J2 Eocercopis ancyloptera Handlirsch, 1939 Germany J1 E. ovalis Handlirsch, 1939 Germany J1 E. similis Popov & Wootton, 1977 Germany J1 Heterojassus membranaceus Evans, 1961 Australia T3 Hexascytina tillyardi Evans, 1956 Australia T3 lIahulgabalus endaidus Szwedo, Azar & Ziadé, 2011 Lebanon K1 Ildavia incompleta Klimaszewski & Popov, 1993 UK K1 I. shcherbakovi Klimaszewski & Popov, 1993 UK K1 Ind...…”

The earliest fossil flower bugs (Heteroptera: Cimicomorpha: Cimicoidea: Vetanthocoridae) from the Middle Jurassic of Inner Mongolia, China

“…Although we did not examine these fossil specimens, based on the available evidence we prefer to treat them as representatives of the Pachymeridiidae. Furthermore, during the Mesozoic, veins on the membrane of Pachymeridiidae diversified (Popov & Wootton, 1977;Yao et al, 2008). So we can't classify the Pachymeridiidae based on some features of the forewing membrane or only one poorly preserved specimen.…”

“…According to the description, illustrations and photographs in the original paper, all the abovementioned species have the following characters: hemelytron divided into a distinct corium and membrane, corium with deep costal fracture, with corium-membrane boundary sub-parallel, cuneus absent, Sc, R and M diverging from a single point. These characters are the main diagnostic features of the Pachymeridiidae, an extinct true bug family that is considered to be ancestral to the Coreoidea (sensu lato, including Coreoidea, Pyrrhocoroidea and Lygaeoidea) and lived during the Mesozoic (Handlirsch, 1906;Popov & Wootton, 1977;Shcherbakov & Popov, 2002;Yao et al, 2008). Although we did not examine these fossil specimens, based on the available evidence we prefer to treat them as representatives of the Pachymeridiidae.…”

New species of Cicadocoris (Hemiptera: Coleorrhyncha: Progonocimicidae) from mid-Jurassic deposits in northeastern China

“…They were widely distributed, occurring in Permian deposits in Australia (Tillyard, 1926), Triassic in Kyrgyzstan (Becker-Migdisova, 1958;Popov & Shcherbakov, 1991), Australia (Evans, 1956(Evans, , 1961(Evans, , 1963Wootton, 1963) and Argentina (Martins-Neto, 2003), Jurassic in Kyrgyzstan (Popov, 1982(Popov, , 1985, United Kingdom (Popov et al, 1994), China (Hong, 1983;Dong et al, 2012Dong et al, , 2013Dong et al, , 2014, Russia (Popov, 1985(Popov, , 1988, Germany (Handlirsch, 1939;Popov & Wootton, 1977), Luxembourg (Szwedo, 2011), and Cretaceous in Lebanon , United Kingdom (Handlirsch, 1906;Klimaszewski & Popov, 1993), Argentina (Martins-Neto, 2003), Mongolia plete or nearly complete, and 16 of them are with counterparts. Most specimens (39) were collected from several localities near Daohugou and Wuhua, Ningcheng County, Inner Mongolia, northeastern China (Fig.…”