A new Progonocimicidae species, Cicadocoris anisomeridis sp.n., with asymmetrical tegmina is described from the Middle Jurassic Jiulongshan Formation in northeastern China. This is the fifth report of Coleorrhyncha from China. A cladistic analysis based on a combination of fossil and extant taxa clarifies the phylogenetic status of the new fossils and allows the reconstruction of inter‐subfamily relationships within the suborder Coleorrhyncha. Coleorrhyncha is monophyletic and divided into two main clades. Progonocimicidae comprises a monophyletic lineage, to which the new fossils belong. The broadly conceived Progonocimicinae and Cicadocorinae, as recognized by earlier authors, are not supported. The monophyly of the family Karabasiidae is also not supported, and its two constituent subfamilies Hoploridiinae and Karabasiinae are raised to family rank. Hoploridiidae is found to be sister group to all extant moss bugs, and Karabasiidae is found to be the monophyletic sister group to Hoploridiidae + all extant moss bugs.
This published work has been registered in ZooBank, http://zoobank.org/urn:lsid:zoobank.org:pub:30BC0498-3B8A-4650-BC9D-02A0C8D870B2.
A new fossil species, Cicadocoris varians sp. n., of the family Progonocimicidae is described from the Middle Jurassic Jiulongshan Formation of Daohugou Village, Inner Mongolia, China. This Progonocimicidae is a well-preserved unusual fossil showing different sizes and proportions of the left and right tegmen.
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