for suggesting this problem, and for his valuable advice and encouragement throughout the investigation. I also wish to thank Dr V. Cosslett for granting me the facilities of the electron microscope atthe Cavendish Laboratory, UniversityofCambridge, and Mr R. Home for taking the electron micrographs. This work was carried out while holding a Travelling Scholarship awarded by the Gowrie Scholarship Trust of Australia. My thanks are also due to the N.S.W. Milk Board for some financial assistance.