The effects of light intensity and nutrient availability on the biomass allocation of Calamagrostis canadensis were studied under both greenhouse and field conditions In the greenhouse, seedlings from forest and wetland populations were grown in sand‐peat mixtures, under three light intensities and three fertilizer levels Total above and below ground growth increased with light intensity and nutrient availability, nutrient availability, however, only altered plant growth when light intensity was above moderate levels Numbers of rhizomes were greatest under low and moderate nutrient regimes but high light was also needed for maximum numbers of rhizomes In the field, tillers and rhizomes were examined under open, 40% and 85% canopy cover Tiller and rhizome weight and diameter of rhizomes decreased with canopy closure However, proportional allocation of biomass to rhizomes was greatest and percentage total non‐structural carbohydrates of rhizomes was lowest under a 40% canopy
Outbreaks of mountain pine beetle are evaluated as a generic disturbance agent, and comparisons are made with other forest disturbances such as wildfire, windthrow, and logging. A useful basis for comparison is the degree of disruption to the overstorey, understorey, and forest floor layers. Clear differences are observed in the impacts of bark beetles, fire, and windthrow, but there is overlap with various harvesting systems. Insects are selective in terms of the species or size of tree that is killed; this selectivity varies with stand composition, stand structure, and outbreak stage. The mountain pine beetle functions as part of larger natural disturbance regimes in western North America, which vary with climate and forest type. Outbreaks of many different insects occur throughout western Canada, with the relative role of fire and insects differing among ecoregions and over time. Beetle-killed stands may facilitate extreme fire behaviour and may be more susceptible to future burning. Large expanses of dead or removed trees also result in altered soil water balance and stream flows, disposing some sites to mass movement or flooding. All disturbances generate heterogeneity, with much of the value to biodiversity and ecosystem recovery depending on residual structure and biological legacies. The capacity for unassisted recovery and the value of each stand to timber supply, carbon balance, and habitat needs in a landscape context are relevant when considering salvage logging or forest rehabilitation. The future role of forest pests is expected to fluctuate in response to changes in climate and the altered composition and structure of western forests.
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