On the cover: Distribution of green alder (Alnus crispa (Ait.) Pursh) stems (light green circles) that occurred in the understory of a mature boreal forest stand at Trapper Creek, Alaska, overlain with postharvest plantation of white spruce (Picea glauca (Moench) Voss) seedlings (darker green symbols). The estimated aboveground biomass of each alder stem is represented by circle diameter, and the size of spruce symbols represents seedling height growth during the first 3 years after planting. The growth of the spruce seedlings was not related to preharvest alder stem locations. In this figure, only the 25 measured seedlings per plot are shown; the plots lacking spruce seedlings entirely are natural regeneration plots. Alders planted with spruce in some of the plots are not shown. To document possible soil nitrogen mosaics before timber harvesting on three boreal forest sites in Alaska, maps of the distribution of understory green (Alnus crispa (Ait.) Pursh) and Sitka alder (A. sitchensis (Reg.) Rydb.) stems were made. Understory alders were regularly distributed throughout the northernmost site (Standard Creek) and very irregularly distributed at the southernmost site (Cooper Landing). No consistent relations existed between alder stem location and total soil nitrogen. In undisturbed forest, soils collected beneath alders tended to have more nitrogen than soils without alder, but after the sites were harvested, soil chemistry differed. To examine the interactions of alder and white spruce (Picea glauca (Moench) Voss) on secondary successional sites, mixed plantations of white spruce and alder were established after each site was harvested. Despite good survival, the planted alder grew poorly. No differences were found between nursery-grown alder seedlings and alder wildlings in either growth or survival. Although fifth-year survival and growth of white spruce were excellent on all sites, they were not related to either the preharvest distribution of naturally occurring alder or to alders planted in the mixed plantations. Locational information and site maps are provided for future evaluation of these plantations.Keywords: White spruce, green alder, Sitka alder, boreal forest, interior Alaska, mixed-species plantations, nitrogen fixation, alder wildlings, long-term ecosystem productivity.Although alders are ecologically important in the boreal forest, their role in managed forest systems has received little attention. This study examined several aspects of the interactions between two shrubby alder species (Alnus crispa (Ait.) Pursh) and A. sitchensis (Reg.) Rydb.) and white spruce (Picea glauca (Moench) Voss), the primary commercial species in the boreal forest of Alaska. First, I mapped the alder stems that occurred naturally in the understory of three mature forest stands and collected soil from beneath alder clumps and from beneath other typical types of vegetation for each site. Soils also were collected after each site was harvested. In undisturbed forest, soils collected beneath alders tended to have more ...