a b s t r a c tHolocene climatic variability and human impact on vegetation are reconstructed from a region in central European Russia, which lies at an important ecotone between broadleaf forest and steppe. For the first time in this region we adopt a multi-proxy approach that combines analysis of local mire conditions from plant macrofossil and testate amoeba analyses with pollen-based quantitative climate reconstruction. The proxies indicate a long-term warming trend from 9700 to 7500 cal yr BP, interrupted by a series of short-term cold events. From 7500 to 5000 cal yr BP the results imply a relatively stable climate, warmer and drier than present, spanning the Holocene Thermal Maximum. Since 5000 cal yr BP the data suggest a change to cooler climate, but with centennial-scale variability. This shift at around 5000 cal yr BP is supported by extensive evidence from other sites. In the early Holocene, the region was occupied mainly by pine and birch forests. Broad-leafed forests of oak, lime and elm expanded after 7800 cal yr BP and remained dominant until the last few centuries. During the historical period, vegetation changes have been driven mainly by human activities.

This paper is focused on the discussion of the exact age, and environmental setting of the 'Khvalynian' Transgression of the Caspian Sea with the use of novel radiometric, isotopic evidence and GIS-linked modelling. Based on this evidence the Khvalynian transgression is viewed as coeval with the warm Late Glacial episodes in the higher latitudes, reaching its peak at 13.6 -11.8 ka. During that period the Caspian Sea was connected with the Caspian Sea via the 'Manych-Kerch-Spillway' north of the Caucasus Mountains. We also suggest a younger age for Mousterian sites in the Caspian basin, which are usually associated with the populations of Neanderthal humans. The initial emergence of Mousterian industries in this area was broadly contemporaneous with MOI 3 that included the LGM (25-18 ka). Their expansion was largely coeval with the Khvalynian transgression.

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The Sukhaya Mechetka is the most important Middle Paleolithic site in the Eastern Europe. The paper presents the analysis of the section, including its cultural layer, in the context of stratigraphy and paleogeography of the Lower Volga River region. The structure of the section represents 11 main stages of sedimentation and paleogeographic evolution of the site territory, which are closely related to global and regional changes in the climate and the level of the Caspian Sea. The base of the section reflects the stage of a river existence on the territory of the site that flowed into the Volga River estuary, formed by the Early Khazarian transgression of the Caspian Sea at the end of the Middle Pleistocene (MIS 6). A long continental period of evolution under multi-directional climate fluctuations of different amplitudes from the Mikulino interglacial (MIS 5e) till the Late Valdai glacial epoch (MIS 2) is reflected in the middle part of the section. The Late Khazarian and Hyrcanian transgressive basins (MIS 5), as well as the Paleo-Khvalynian stage (MIS 3) of the Caspian Sea did not reach the latitude of Sukhaya Mechetka. The stage of Early Khvalynian transgression (MIS 2, 16 190 ± 200 years ago) during the degradation of Ostashkovo glaciation is expressed in the upper part of the section. Three paleosoils have been found that reflect the warming periods of the MIS 5 stage and are related to its 5e, 5c and 5a sub-stages. The middle paleosoil contains a cultural layer assigned to the Micoquian/KMG community of Northern Eurasia. The climate was moderately warm during the Neanderthal settlement of the territory, steppe landscapes dominated the interfluve, and the forests grew in the balka. The Hyrcanian transgression of the Caspian Sea with an estuary in the Volga River valley predetermined a high erosion basis and the formation of a balka with a wide bottom and gentle banks. A permanent fresh watercourse attracted mammals which were an object of hunting for ancient men. All this, obviously, became a basis for the organization of settlement in the Sukhaya Mechetka balka. Correlation of the section with detailed studied loess-soil sections of the Lower Volga River region makes it possible to determine the age of its cultural horizon in between 97-110 thousand years ago.