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A recent collection of actinopterygian fossil fishes from a previously unreported locality in the Cenomanian or Turonian of southeastern Morocco includes a single specimen of a macrosemiid fish. Macrosemiids are more common in Jurassic and Early Cretaceous deposits, with the previously known range of the family being Late Triassic through Aptian or Albian. This discovery therefore extends the temporal range of the family into the Late Cretaceous. Moreover, macrosemiids had not previously been reported from northern Africa or the Moroccan area of the Tethys basin; therefore, this fossil also increases the geographical range of the family. The Moroccan macrosemiid is described in a new genus and species, Agoultichthys chattertoni. A phylogenetic analysis places it basal to all other genera of the family with the exception of Notagogus. Diagnostic characters of the new species include the high number of scales laterally along the body and the greater number of dorsal fin rays than in other members of the family.
A euselachian assemblage was recovered from the middle Cenomanian Dunvegan Formation situated in northwestern Alberta. This assemblage is the most northern described within the Western Interior Seaway and provides an important insight into the euselachian faunal diversity of this little-known region of the seaway. Despite its high paleolatitude, the assemblage contains a number of elasmobranch taxa, including Hybodus , Squalicorax , Archaeolamna , Cretodus , Dallasiella , and Cretoxyrhina . The Dunvegan assemblage also contains the first known reports from Canada of the odontaspid shark Johnlongia parvidens , the cretoxyrhinid shark Protolamna carteri , and the ray Pseudohypolophus mcnultyi . This assemblage extends the northern geographical range of all taxa. Preliminary comparisons with other middle Cenomanian Western Interior Seaway assemblages show that the core composition of the Dunvegan assemblage is remarkably similar to that of other time-equivalent assemblages; however, conspicuously absent are species that are exceedingly common in other localities situated farther south. We suggest that absence of these taxa from the Dunvegan localities may be caused by a temperature intolerance associated with latitude and sea-water circulation patterns, or by an inability to inhabit environments that exhibit salinity variation.
A new genus and species of osteoglossomorph fish is described from the Eocene Mahenge site in north central Tanzania. The new species is sympatric with the previously described osteoglossomorph † Singida jacksonoides , but is easily distinguished from it by the presence in the new species of teeth on the jaws and entopterygoid. The new species shares characters with the Heterotidinae, such as the two arms of the preopercle being of similar length and the posterior end of the maxilla lying on the dentary, and other characters with the Osteoglossinae (including † Phareodus ), for example the enlarged pectoral fin ray and the shape of the opercular bone. New specimens of † S. jacksonoides also have been recovered from the site, providing more information on the osteology of this fish. The relationships of these two Mahenge osteoglossomorphs are assessed, with both fishes placed within the Osteoglossidae, but of uncertain position in the family. Addition of the new species to previous cladistic data sets, and the revised information for † S. jacksonoides , causes the subfamily Osteoglossinae to be nonmonophyletic.forward almost to the level of the midpoint of the orbit, articulation of the anguloarticular and quadrate just posterior to the level of the midpoint of the orbit, the angle formed by the maxilla and dentary anterior to the level of the midpoint of the orbit, and the posterior end of the maxilla lying on the dentary. Distinguished from † Brychaetus, † Phareodus and † Cretophareodus by the lack of reticulate scales in the new species; from † Sinoglossus, Heterotis, Arapaima, † Laeliichthys and † Joffrichthys by the presence of an enlarged, long
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