el; of Silurian soils as a result of pedoturbat i o~~, effectively mcreasing the average depth of soil CO, proiluction.Our results (Table 1) ~m p l y that atmospheric CO-, decllneil 1~y a factor of 10 from the Late S i h r i a n to the Early Permian, closely follow~ng (Fig. 4 ) a decline precllctecl hi; theoretical carbon lnais balance models (1). T h e largest decrease, hetween the Late Sil~lrian a11il Late Devonian. coincides with a of rapid evolution and diversificatlon of the terrestrial ecosystem (18).Estimates of atmospheric C02 levels from separated, time-equivalent ~-7aleosols are consistent, suggertlng that a coherent record of changing atlnospheric chem~stry is yreserl-eii In the ancient soil recorJ. REFERENCES AND NOTES1 R A Berner Science 261, 68 (1 993) At?? J SCI 294 56 (1 994) 2 T J Crowley and G R North, Paleoc!~~rato!ogy (Oxford UI?I\/ Press, Oxford, 1991) 3 R A Berner and R Ralswell, Geochim. Cosmochlm. Acta 47. 855 11983) L R. I

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