Two-WingedCloeodesin Brazil: New Species, Stage Description, and Key to South American Species

Abstract

The present work, based on material from northern, central-western, and northeastern Brazil, contributes to the knowledge of the two-winged Cloeodes Traver (Ephemeroptera: Baetidae) in South America. Two new species, C. maracatu, sp. nov. and C. spaceki, sp. nov., are described, the former based on nymphs and reared adults and the latter only on nymphs; the male and female imago of C. auwe and the female imago of C. redactus are described. Based on these findings, an updated key for South American nymphs and male adults of the two-winged Cloeodes is provided.

Phylogenetic analyses of CloeodesTraver and related genera (Ephemeroptera: Baetidae)

“…One of the species of Cloeodes described by Lugo‐Ortiz & McCafferty () from Australia was revalidated and transferred to Bungona Harker (Webb & Suter, ). Finally, the number of species of Cloeodes doubled in the Neotropics during this period, with more than ten species described from Argentina, Brazil and Uruguay (Salles & Lugo‐Ortiz, ; Salles et al ., ; Nieto & Richard, ; Massariol & Salles, ; Nieto & Emmerich, ; Massariol et al ., ).…”

New species and records of Cloeodes Traver, 1938 (Ephemeroptera, Baetidae) from Costa Rica

“…Individuals were found at 2054 m asl in cold waters of two cloud forest streams (Fig. 7); other non-Andean species have been reported between 159-1800 m asl (Nieto and Richard 2008;Gutiérrez and Reinoso-Flórez 2010;Massariol and Salles 2011;Massariol et al 2013;De Paul et al 2013;Gutiérrez and Dias 2015). Also, the new species was collected in a pristine forest region, which may be similar to other Cloeodes species in Brazil that seem to prefer remnant forest areas (Salles et al 2015).…”

Baetidae (Insecta: Ephemeroptera) from the state of Rondônia, Northern Brazil

“…Brazil: Mato Grosso (Salles et al 2004b); Roraima (Falcão et al 2012); Rondônia (Massariol et al 2013).…”