“…While the addition of mineral fertilizer only increases nutrient availability, the addition of organic fertilizer can also increase ion-exchange capacity (reducing nutrient leaching) and water retention capacity (Johnson, 1987;Elmarsdottir et al, 2003). Regeneration of indigenous plant cover can also be inhibited by factors common to all regions, including the absence of viable seeds (Ebersole, 1989;Forbes and Jefferies, 1999;Prach et al, 2001) or of mycorrhizal inoculum (Greipsson and El-Mayas, 1999;Blanke et al, 2007), herbivory (Arnalds, 1987;Bradshaw, 1987), substrate instability (Walker and Del Moral, 2003), compaction (Billings, 1987), desiccation (Arnalds, 1987;Cargill and Chapin, 1987;Chapin, 1993), and low nutrient availability (Arnalds, 1987;Bradshaw, 1987;Blanke et al, 2007).…”