is an essential nutrient for all living organisms which enters the food chain through its acquisition from environmental sources by primary producers. In approximately 50% of agricultural soils, P is limiting for plant growth and supplemental fertilization is required for optimal crop productivity (Lynch, 2011). However, P-rich rock required for inorganic phosphate fertilizer production is a limited, nonrenewable resource (Elser & Bennett, 2011; Steiner, Geissler, Geissler, Watson, & Mew, 2015). Paradoxically, a high proportion of P applied as fertilizer is not available to plants due to its fixation in the soil by chemical and biological processes (Bieleski,