“…The measured reflectance spectra are sensitive to the structural organization of, and variations in chemical constituents in, canopy components. These physico-chemical-tospectral linkages provide a means of detecting species and/or functional types (e.g., Asner & Vitousek, 2005;Clark, Roberts, & Clark, 2005;Somers & Asner, 2012;Ustin & Gamon, 2010), and can even provide information about the biogeochemical heterogeneity (e.g. Townsend, Asner, & Cleveland, 2008;Vitousek, Asner, Chadwick, & Hotchkiss, 2009) and species richness of tropical forest canopies (e.g., Asner, Nepstad, Cardinot, & Ray, 2004;Carlson, Asner, Hughes, Ostertag, & Martin, 2007;Feret & Asner, 2013;Kalacska et al, 2007;Nagendra & Rocchini, 2008;Somers and Asner, 2013).…”