“…This reduction in shoot length and needle lifespan in both pines at the damaged site is the result of photosynthetic suppression stemming from environmental stressors such as F and excess Mn and Cl. Needle loss and premature senescence due to air pollutants such as SO 2 , O 3 , NO x , or acid deposition, have been reported in pine, birch and poplar (Reich, 1983;Reich et al, 1995; The toxicity of these pollutants is mainly due to their interference with respiration and photosynthetic function (Heber et al, 1995;Lambers et al, 1998;Matyssek et al, 1995a, b;Pukacki, 2000). The energy status of damaged trees may then be reduced (Ernst, 1976;Ernst and Joosse-van Damme, 1983) by smaller uptake of mineral nutrients (Adams, 1981;Larcher, 2003).…”