Plant traits – the morphological, anatomical, physiological, biochemical and phenological characteristics of plants and their organs – determine how primary producers respond to environmental factors, affect other trophic levels, influence ecosystem processes and services and provide a link from species richness to ecosystem functional diversity. Trait data thus represent the raw material for a wide range of research from evolutionary biology, community and functional ecology to biogeography. Here we present the global database initiative named TRY, which has united a wide range of the plant trait research community worldwide and gained an unprecedented buy-in of trait data: so far 93 trait databases have been contributed. The data repository currently contains almost three million trait entries for 69 000 out of the world's 300 000 plant species, with a focus on 52 groups of traits characterizing the vegetative and regeneration stages of the plant life cycle, including growth, dispersal, establishment and persistence. A first data analysis shows that most plant traits are approximately log-normally distributed, with widely differing ranges of variation across traits. Most trait variation is between species (interspecific), but significant intraspecific variation is also documented, up to 40% of the overall variation. Plant functional types (PFTs), as commonly used in vegetation models, capture a substantial fraction of the observed variation – but for several traits most variation occurs within PFTs, up to 75% of the overall variation. In the context of vegetation models these traits would better be represented by state variables rather than fixed parameter values. The improved availability of plant trait data in the unified global database is expected to support a paradigm shift from species to trait-based ecology, offer new opportunities for synthetic plant trait research and enable a more realistic and empirically grounded representation of terrestrial vegetation in Earth system models.

In preparation for the fifth Assessment Report (AR5) of the Intergovernmental Panel on Climate Change (IPCC), the international community is developing new advanced Earth System Models (ESMs) to assess the combined effects of human activities (e.g. land use and fossil fuel emissions) on the carbon-climate system. In addition, four Representative Concentration Pathway (RCP) scenarios of the future (Moss et al. 2010). The diversity of approaches and requirements among IAMs and ESMs for tracking land-use change, along with the dependence of model projections on land-use history, presents a challenge for effectively passing data between these communities and for smoothly transitioning from the historical estimates to future projections. Here, a harmonized set of land-use scenarios are presented that smoothly connects historical reconstructions of land use with future projections, in the format required by ESMs. The land-use harmonization strategy estimates fractional land-use patterns and underlying land-use transitions annually for the time period 1500-2100 at 0.5°×0.5°resolution. Inputs include new gridded historical maps of crop and pasture data from HYDE 3.1 for 1500-2005, updated estimates of historical national wood harvest and of shifting cultivation, and future information on crop, pasture, and wood harvest from the IAM implementations of the RCPs for the period 2005-2100. The computational method integrates these multiple data sources, while minimizing differences at the transition between the historical reconstruction ending conditions and IAM initial conditions, and working to preserve the future changes depicted by the IAMs at the grid cell level. This study for the first time harmonizes land-use history data together with future scenario information from multiple IAMs into a single consistent, spatially gridded, set of land-use change scenarios for studies of human impacts on the past, present, and future Earth system.

The Community Land Model is the land component of the Community Climate System Model. Here, we describe a broad set of model improvements and additions that have been provided through the CLM development community to create CLM4. The model is extended with a carbon‐nitrogen (CN) biogeochemical model that is prognostic with respect to vegetation, litter, and soil carbon and nitrogen states and vegetation phenology. An urban canyon model is added and a transient land cover and land use change (LCLUC) capability, including wood harvest, is introduced, enabling study of historic and future LCLUC on energy, water, momentum, carbon, and nitrogen fluxes. The hydrology scheme is modified with a revised numerical solution of the Richards equation and a revised ground evaporation parameterization that accounts for litter and within‐canopy stability. The new snow model incorporates the SNow and Ice Aerosol Radiation model (SNICAR) ‐ which includes aerosol deposition, grain‐size dependent snow aging, and vertically‐resolved snowpack heating – as well as new snow cover and snow burial fraction parameterizations. The thermal and hydrologic properties of organic soil are accounted for and the ground column is extended to ∼50‐m depth. Several other minor modifications to the land surface types dataset, grass and crop optical properties, surface layer thickness, roughness length and displacement height, and the disposition of snow‐capped runoff are also incorporated. The new model exhibits higher snow cover, cooler soil temperatures in organic‐rich soils, greater global river discharge, and lower albedos over forests and grasslands, all of which are improvements compared to CLM3.5. When CLM4 is run with CN, the mean biogeophysical simulation is degraded because the vegetation structure is prognostic rather than prescribed, though running in this mode also allows more complex terrestrial interactions with climate and climate change.

Abstract. Regional phenology is important in ecosystem simulation models and coupled biosphere/atmosphere models. In the continental United States, the timing of the onset of greenness in the spring (leaf expansion, grass green-up) and offset of greenness in the fall (leaf abscission, cessation of height growth, grass brown-off) are strongly influenced by meteorological and climatological conditions. We developed predictive phenology models based on traditional phenology research using commonly available meteorological and climatological data. Predictions were compared with satellite phenology observations at numerous 20 km x 20 km contiguous landcover sites. Onset mean absolute error was 7.2 days in the deciduous broadleaf forest (DBF) biome and 6.1 days in the grassland biome. Offset mean absolute error was 5.3 days in the DBF biorne and 6.3 days in the grassland biome. Maximum expected errors at a 95% probability level ranged from 10 to 14 days. Onset was strongly associated with temperature summations in both grassland and DBF biomes; DBF offset was best predicted with a photoperiod function, while grassland offset required a combination of precipitation and temperature controls.

A method for generating daily surfaces of temperature, precipitation, humidity, and radiation over large regions of complex terrain is presented. Required inputs include digital elevation data and observations of maximum temperature, minimum temperature and precipitation from ground-based meteorological stations. Our method is based on the spatial convolution of a truncated Gaussian weighting filter with the set of station locations. Sensitivity to the typical heterogeneous distribution of stations in complex terrain is accomplished with an iterative station density algorithm. Spatially and temporally explicit empirical analyses of the relationships of temperature and precipitation to elevation were performed, and the characteristic spatial and temporal scales of these relationships were explored. A daily precipitation occurrence algorithm is introduced, as a precursor to the prediction of daily precipitation amount. Surfaces of humidity (vapor pressure deficit) are generated as a function of the predicted daily minimum temperature and the predicted daily average daylight temperature. Daily surfaces of incident solar radiation are generated as a function of Sun-slope geometry and interpolated diurnal temperature range. The application of these methods is demonstrated over an area of approximately 400000 km 2 in the northwestern USA, for I year, including a detailed illustration of the parameterization process. A cross-validation analysis was performed, comparing predicted and observed daily and annual average values. Mean absolute errors (MAE) for predicted annual average maximum and minimum temperature were 0.7°C and 1.2°C, with biases of +0. loC and -0. loC, respectively. MAE for predicted annual total precipitation was 13.4 cm, or, expressed as a percentage of the observed annual totals, 19.3%. The success rate for predictions of daily precipitation occurrence was 83.3%. Particular attention was given to the predicted and observed relationships between precipitation frequency and intensity, and they were shown to be similar. We tested the sensitivity of these methods to prediction grid-point spacing, and found that areal averages were unchanged for grids ranging in spacing from 500 m to 32 km. We tested the dependence of the results on timestep, and found that the temperature prediction algorithms scale perfectly in this respect. Temporal scaling of precipitation predictions was complicated by the daily * Corresponding author.0022-1694/97/$17.00 @ 1997-Elsevier Science B.V. All rights reserved PIl SOO22-1694(96)03128-9Journal of P.E. Thornton et al./Journal of Hydrology 190 ( 1997) 214-251 215 occurrence predictions, but very nearly the same predictions were obtained at daily and annual timesteps. @ 1997 Elsevier Science B.V.