The olfactory systems of insects can detect semiochemicals and transfer signals in complex environments, which can initiate specific behaviours in insects, such as searching for food, locating hosts, finding mates, selecting oviposition sites and avoiding enemies (Leal, 2013;Takken & Knols, 1999). Various olfactory proteins are involved in the chemoreceptive process in antennal sensilla, including odorant-binding proteins (OBPs), chemosensory proteins, odorant receptors, ionotropic receptors, sensory neuron membrane proteins and odorant-degrading enzymes (Vogt, 2003;Zhao et al., 2017;Zhou et al., 2004). Insect OBPs are small and water-soluble proteins that can transport specific chemicals to membrane-bound olfactory receptors (Brito et al., 2016). Based on amino acid sequences and expression patterns, lepidopteran OBPs are usually divided into three subfamilies: general odorant-binding proteins (GOBPs), pheromonebinding proteins and antennal binding proteins (Breer et al., 1990;
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