The mid-European segment of the Variscides is a tectonic collage consisting of (from north to south): Avalonia, a Silurian-early Devonian magmatic arc, members of the Armorican Terrane Assemblage (ATA: Franconia, Saxo-Thuringia, Bohemia) and Moldanubia (another member of the ATA or part of N Gondwana?).The evolution on the northern flank of the Variscides is complex. Narrowing of the Rheic Ocean between Avalonia and the ATA occurred during the late Ordovician through early Emsian, and was accompanied by formation of an oceanic island arc. By the early Emsian, the passive margin of Avalonia, the island arc and some northern part of the ATA were closely juxtaposed, but there is no tectonometamorphic evidence of collision. Renewed extension in late Emsian time created the narrow Rheno-Hercynian Ocean whose trace is preserved in South Cornwall and at the southern margins of the Rhenish Massif and Harz Mts. Opening of this ‘successor ocean’ to the Rheic left Armorican fragments stranded on the northern shore. These were later carried at the base of thrust sheets over the Avalonian foreland. Closure of the Rheno-Hercynian Ocean in earliest Carboniferous time was followed by deformation of the foreland sequences during the late lower Carboniferous to Westphalian.Closure of narrow oceanic realms on both sides of Bohemia occurred during the mid- and late Devonian by bilateral subduction under the Bohemian microplate. In both these belts (Saxo-Thuringian, Moldanubian), continental lithosphere was subducted to asthenospheric depths, and later partially obducted. Collisional deformation and metamorphism were active from the late Devonian to the late lower Carboniferous in a regime of dextral transpression. The orthogonal component of intra-continental shortening produced an anti-parallel pair of lithospheric mantle slabs which probably joined under the zone of structural parting and became detached. This allowed the ascent of asthenospheric material, with important thermal and rheological consequences. The strike slip displacements were probably in the order of hundreds of kilometres, since they have excised significant palaeogeographic elements.

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The Montagne Noire forms the southernmost part of the French Massif Central. Carboniferous flysch sediments and very low-grade metamorphic imprint testify to a very external position in the orogen. Sedimentation of synorogenic clastic sediments continued up to the Viséan/Namurian boundary (B320 Ma). Subsequently, the Palaeozoic sedimentary pile underwent recumbent folding and grossly southward thrusting. An extensional window exposes a hot core of Carboniferous HT/LP gneisses, migmatites and granites (Zone Axiale), which was uplifted from under the nappe pile. After the emplacement of the nappes on the Zone Axiale (Variscan D 1 ), all structural levels shared the same tectonic evolution: D 2 (extension and exhumation), D 3 (refolding) and post-D 3 dextral transtension. HT/LP-metamorphism in the crystalline rocks probably started before and continued after the emplacement of the nappes. Peak metamorphic temperatures were attained during a post-nappe thermal increment (M 2 ). M 2 occurred during ENE-directed bilateral extension, which exhumed the Zone Axiale and its frame as a ductile horst structure, flanked to the ENE by a Stephanian intra-montane basin. Map patterns and mesoscopic structures reveal that extension in ENE occurred simultaneously with NNW-oriented shortening. Combination of these D 2 effects defines a bulk prolate strain in a ''pinched pull-apart'' setting. Ductile D 2 deformation during M 2 dominates the structural record. In wide parts of the nappes on the southern flank of the Zone Axiale, D 1 is only represented by the inverted position of bedding (overturned limbs of recumbent D 1 folds) and by refolded D 1 folds. U-Pb monazite and zircon ages and K-Ar muscovite ages are in accord with Ar-Ar data from the literature. HT/LP metamorphism and granitoid intrusion commenced already at C330 Ma and continued until 297 Ma, and probably in a separate pulse in post-Stephanian time. Metamorphic ages older than c. 300 Ma are not compatible with the classical model of thermal relaxation after stacking, since they either pre-date or too closely post-date the end of flysch sedimentation. We therefore propose that migmatization and granite melt generation were independent from crustal thickening and caused, instead, by the repeated intrusion of melts into a crustal-scale strike-slip shear zone. Advective heating continued in a pull-apart setting whose activity outlasted the emplacement of the Variscan nappe pile. The shearzone model is confirmed by similar orogen-parallel extensional windows with HT/LP metamorphism and granitoid intrusion in neighbouring areas, whose location is independent from their position in the orogen. We propose that heat transfer from the mantle occurred in dextral strike-slip shear zones controlled by the westward