Araucarioxylon Kraus is a widely known fossil-genus generally applied to woods similar to that of the extant Araucariaceae. However, since 1905, several researchers have pointed out that this name is an illegitimate junior nomenclatural synonym. At least four generic names are in current use for fossil wood of this type: Agathoxylon Hartig, Araucarioxylon, Dadoxylon Endl. and Dammaroxylon J.Schultze-Motel. This problem of inconsistent nomenclatural application is compounded by the fact that woods of this type represent a wide range of plants including basal pteridosperms, cordaitaleans, glossopterids, primitive conifers, and araucarian conifers, with a fossil record that extends from the Devonian to Holocene. Conservation of Araucarioxylon has been repeatedly suggested but never officially proposed. Since general use is a strong argument for conservation, a poll was conducted amongst fossil wood anatomists in order to canvass current and preferred usage. It was found that the community is divided, with about one-fifth recommending retention of the well-known Araucarioxylon, whereas the majority of others advocated use of the legitimate Agathoxylon. The arguments of the various colleagues who answered the poll are synthesized and discussed. There is clearly little support for conservation of Araucarioxylon. A secondary aspect of the poll tackled the issue as to whether Araucaria-like fossil woods should be either gathered into a unique fossil-genus, or whether two fossil-genera should be recognized, based on the respective presence or absence of axial parenchyma. A majority of colleagues favoured having one fossil-genus only. Agathoxylon can be used legitimately and appears to be the most appropriate name for such woods. However, its original diagnosis must be expanded if those woods lacking axial parenchyma are to be included.
Abstract.A new fossil conifer wood -Brachyoxylon currumilii sp. nov.-is described from the Lower-Middle Jurassic of Chubut Province (Argentina). The specimens were collected at a locality where the Cañadón Asfalto Formation is exposed, in the vicinity of the Cerro Cóndor village. The studied wood is characterized by mixed pitting in radial tracheid walls (predominantly uniseriate), araucarioid cross fields, low uniseriate rays and the absence of resin canals. The use of transmitted light and epifluorescence microscopy together with SEM imaging allowed the identification of additional characters such as the presence of pitting with torus-margo and compression wood. These characters are discussed in terms of their systematic relevance. On these grounds, the systematic affinity of the studied wood is discussed and linked to the extinct conifer family Cheirolepidiaceae, one of the dominant plant groups during the Jurassic and Cretaceous in Patagonia.Keywords. Fossil wood. Mesozoic. Patagonia. Brachyoxylon. Cheirolepidiaceae.
Resumen. PRIMER REGISTRO DE LEÑO DE CONIFERA DE LA FORMACIÓN CAÑADÓN ASFALTO (JURÁSICO TEMPRA-NO-MEDIO), PROVINCIA DE CHUBUT, ARGENTINA. Se describe una nueva especie de madera fósil de conífera, Brachyoxylon currumilii sp. nov., del Jurásico Inferior a Medio de la provincia de Chubut, Argentina. Los especímenes estudiados fueron coleccionados en una localidad donde aflora la Formación Cañadón Asfalto, en las cercanías de la aldea de Cerro Cóndor. Los troncos, estudiados, se caracterizan por el punteado radial de las traqueidas de tipo mixto, predominantemente uniseriado, campos de cruzamiento de tipo araucarioide, radios uniseriados bajos y ausencia de canales resiníferos. La observación bajo MEB y microscopio con luz transmitida y epifluorescencia permitió además la identificación de caracteres tales como la presencia de torus-margo en las punteaduras y estructuras de leño de compresión. Ambos caracteres son discutidos en relación a su relevancia sistemática. Sobre esta base, se discute la afinidad sistemática de la madera estudiada, la cual posee rasgos que la vincularían con la familia de coníferas extintas Cheirolepidiaceae, uno de los grupos de plantas dominantes durante el Jurásico y Cretácico de Patagonia.Palabras clave. Madera fósil. Mesozoico. Patagonia. Brachyoxylon. Cheirolepidiaceae.
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