The Nebraska Sand Hills are a stabilized dune field in the central United States that reflect past conditions of drought. The most recent drought, known as the Medieval Climatic Anomaly, occurred from A.D. 900 to A.D. 1300 and had an enormous effect on the thriving prairie ecosystem, which included large populations of the plains pocket gopher (Geomys bursarius). Burrows of these organisms across a paleosol-eolian sand boundary in the Sand Hills indicate abrupt climate change during the transition from stabilized to active dune field and from humid to arid conditions. Medieval gophers tunneled at greater depths below the surface than do modern gophers, indicating the behavioral changes these animals underwent to survive during the transition. The gophers were likely surviving on roots remaining in the underlying soil as it was buried by sand; they tunneled .1 m up to the surface to deposit mounds of excavated soil and sand. Most of the burrows occur in areas of low-angle bedding, suggesting loss of vegetation occurred first on the crests of the newly formed dunes while vegetation persisted in the interdunes. Optically stimulated luminescence dates from a dune containing ancient gopher burrows are nearly identical throughout the height of the dune, indicating rapid accumulation of sand. As accumulation of sand was rapid, vegetative loss must also have occurred quickly, though not in a uniform pattern across the region. Pocket gophers were apparently able to survive in areas of remaining vegetation for a short time, but in a relatively short period of time, they were unable to reach their food sources and were forced ultimately to abandon the uplands in the region.
Paleogastroliths, as defined in this study, are stomach stones associated exclusively with extinct animals. Definitive recognition of paleogastroliths is problematic and controversial. Polished, rounded stones found within skeletal remains of dinosaurs and plesiosaurs are the strongest evidence. This study examined intraskeletal and proposed paleogastroliths using a scanning electron microscope (SEM) to determine if any defining surface textures occur. Paleogastroliths associated with the skeletal remains of Seismosaurus hallorum, possible paleogastroliths from the Lower Cretaceous Cedar Mountain Formation, and other well-polished stones were obtained. Most stones were composed of either quartzite or chert and were examined with a SEM at magnifications ranging from 100× to 400×. All of the cherty Seismosaurus hallorum stones contained sets of parallel to sub-parallel, curved polish grooves that in places crosscut one another. These grooves are inferred to have formed as the result of small particles lodged in the stomach muscles interacting with the stones due to muscle movement. The distinctive surface textures noted on the cherty Seismosaurus hallorum paleogastroliths are similar to those seen on the cherty possible paleogastroliths from the Cedar Mountain Formation. Therefore, the geologic occurrence combined with the microscopic surface texture supports the interpretation that these stones are paleogastroliths.
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