a b s t r a c tFossil palm leaves from the Oligocene deposits in Ningming County, Guangxi, China, have been reported for the first time. The palmate leaf shape along with a definite costa support the placement of the fossils in the Coryphoideae. Three new species are described: Chuniophoenix slenderifolia sp. nov., Livistona roundifolia sp. nov. and Trachycarpus formosa sp. nov. The palm fossils belong to three taxa and indicate that palms in southern China began to diversify no later than the Oligocene and that a diversified palm flora was already present at that time in Guangxi. This report provides new information about the biogeographical history of Chinese palms over geological time.
Paliurus favonii Unger is recognized and described based on fruits from the Oligocene Ningming flora of Guangxi, South China. Characteristics of the present specimens include circular winged fruits that are 10.0–11.5 mm in diameter with a central endocarp at 3.0 to 4.0 mm in diameter. The specimens fall into the morphological range of the fossil species P. favonii, which has been observed in other Cenozoic sites in the Northern Hemisphere. The present discovery represents the lowest latitude distribution of P. favonii in the world, and we are presenting the first P. favonii fossil described with detailed cuticular characteristics from China. Further, this finding demonstrates that the genus existed in the Oligocene Ningming region, South China, and provides new information for understanding the fossil history. The dispersal mode for winged fossils demonstrates that wind dispersal is well-represented in the Oligocene Ningming flora.
Cupressinocladus Seward is a fossil genus of conifers and conifer fossils with reproductive organs are very rare. In general, it is difficult to understand the natural affinities with other conifers. In this paper, a new species, Cupressinocladus guyangensis P.H. Jin et B.N. Sun sp. nov., is reported based on branches with immature female cones from the Lower Cretaceous Guyang Formation of the Guyang Basin in Inner Mongolia, northern China. The foliage shoots are decussate. Leaves are decussate, imbricate, scale‐like, weakly dimorphic, and bear longitudinal glands on the abaxial view. Stomata complexes are haplocheilic, monocyclic, irregularly arranged, and spread along the leaf margin. Immature female cones are subglobose with 6–8 cone scales, and three subglobose ovules arranged in a row at the base of the cone scales. Moreover, we performed cluster analysis using a statistics and machine learning toolbox for 23 fossils and extant species based on 16 morphological characters. The result implies that the new species bears a close resemblance to the extant Cupressus funebris Endl. and might have nearest systematic affinities to it.
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