Plant functional types (PFT) have been used to describe the response of native vegetation to environmental factors (i.e., fertility) and to livestock disturbance, but rarely under conditions of continuous grazing. In this work we investigate whether the longterm response of grassland communities submitted to a gradient of continuous grazing pressure can be described with such an approach. After 15 yr of differentiation of the grazing pressure applied to native grasslands we measured leaf dry-matter content (LDMC) and specific leaf area (SLA) of Poaceae populations of the communities. A grazing pressure gradient was created by levels of daily forage allowance: 4, 8, 12, and 16 kg of dry matter per day per 100 kg of animal live weight, monitored monthly. PFTs were defined by numerical analysis, where an algorithm finds the optimal trait subset based on the agreement between matrices of species 3 traits, paddocks 3 grass biomass, and environmental variables (levels of forage allowance and soil characteristics). The results show that it is possible to describe a gradient of grazing pressure by means of LDMC and/or SLA measured only on the Poacea contributing at least 80% of the total Poaceae biomass. Four PFTs were differentiated by these leaf traits. PFTs having low LDMC and high SLA are characteristic of high intensity of use and are made up largely of stoloniferous C 4 species typical of rapid resource capture strategies. Conversely, PFTs characterized by high LDMC and low SLA include species that are representative of low grazing pressure. Variations in the aggregate value of traits are due to changes in the species proportions and not to leaf-size adaptation as hypothesized. We conclude than in the absence of a gradient of fertility, plants with strategies of resource capture tend to be more represented under high grazing pressures. This situation results in a loss of functional diversity, but in particular a reduction in forage availability, which is incompatible with high animal production. Resumen La noción de grupos funcionales de plantas (GFP) ha sido utilizada para describir la respuesta de la vegetación nativa a los factores del medio (fertilidad) y al disturbio del pastoreo, pero pocos de dichos estudios han sido conducidos bajo condiciones de pastoreo continuo. En el presente trabajo se trata de verificar si dicho enfoque puede utilizarse para analizar la respuesta de la comunidad a un gradiente de presión de pastoreo continuo. Al te´rmino de 15 añ os de diferenciación continua de la presión de pastoreo ejercida sobre una pradera nativa, se midió el contenido de materia seca foliar (CMSF) y el área foliar específica (AFE) de las poblaciones de gramíneas presentes en la comunidad. El gradiente de pastoreo fue establecido según los niveles de oferta de forraje: 4, 8, 12, y 16 kg de materia seca por 100 kg de peso vivo por día, oferta ajustada mensualmente. Los GFP fueron definidos según un análisis numérico, donde un algoritmo identifica un subgrupo óptimo de características basado en la correspondencia e...

To maximize herbage DMI, pregrazing sward surface height (SSH) and level of herbage depletion (HD) must be such that variables determining short-term herbage intake such as bite mass (BM) and bite rate (BR) are optimized. The objective of this study was to determine a SSH target and the level of HD as a proportion of the SSH that optimizes BM and BR of beef heifers grazing Sorghum bicolor swards. Two experiments were conducted using 2 S. bicolor swards and 4 beef heifers (25 mo old; 322 kg BW). Experiment 1 compared the effect of 6 pregrazing SSH, 30, 40, 50, 60, 70, and 80 cm, on BM, BR, and jaw movements. Experiment 2 assessed the effect of HD level as a proportion of SSH (0.17, 0.34, 0.50, 0.67 and 0.84) on BM, BR, and jaw movements using the optimal pregrazing SSH defined in Exp. 1. Short-term herbage DMI was estimated using a double-weighing technique and corrected for insensible BW loss. Herbage DMI was subsequently used to calculate the BM. Net eating time and jaw movements for apprehension and manipulation + mastication during grazing as well as total jaw movements were determined using the IGER (Institute of Grassland and Environmental Research) behavior recorders. Bite rate and the number of total jaw movements per gram herbage DMI were derived from jaw movement count and measurements of herbage DMI. The results of Exp. 1 showed low and high SSH constraint the ease of herbage harvesting. Greater BM are maintained until a SSH of 50 cm is reached (P < 0.05) and then decline at greater SSH due to herbage dispersion. The nonbiting jaw movement rate increased at greater SSH whereas BR decreased (P < 0.05). For both variables, the turning point was close to a SSH of 50 cm. Experiment 2 showed that such an optimization of BM and BR was maintained until an HD level of 0.34 was reached (P < 0.05). There was a linear increase in both the total jaw movements per unit herbage DMI and the nonbiting jaw movements rate (manipulation + mastication) subsequent to levels of HD greater than 0.34 (P < 0.05). These studies provide, for the first time, sward feature targets to manage grazing and optimize BM and BR, aiming to maximize the short-term herbage DMI of cattle grazing S. bicolor swards.

RESUMO -A utilização de cereais de inverno no sistema de duplo propósito permite fornecer aos animais forragem verde no período crítico de carência alimentar, além de aumentar a estabilidade da receita da produção pela melhoria na qualidade e produtividade dos grãos dos cereais de inverno. O experimento foi conduzido no período de abril de 1994 a setembro de 1996 em Guarapuava, Paraná, a fim de avaliar o potencial de utilização para forragem e grãos de aveia branca (Avena sativa L.), trigo (Triticum aestivum), triticale (X. Triticosecale Witt.), aveia preta (Avena strigosa Schreb), centeio (Secale cereale L.) e cevada (Hordeum vulgare L.), visando sua utilização em condições de duplo propósito. Utilizou-se o delineamento experimental de blocos ao acaso, com tratamentos distribuídos em parcelas subdivididas, em três repetições. Nas parcelas foram estudados os sistemas de produção (sem corte, um e dois cortes) e nas subparcelas, os genótipos. O sistema de dois cortes foi superior aos demais quanto ao rendimento de matéria seca, principalmente para a aveia. Para o rendimento de grãos, os sistemas sem corte e um corte foram superiores, apesar da maior produção dos genótipos de aveia sob dois cortes. Em todos os genótipos, houve melhoria do peso do hectolitro e redução da massa de mil sementes, quando se realizaram cortes. Sob condições de manejo adequadas, pode-se alcançar considerável produção de forragem, sem afetar a posterior produção de grãos para cereais de inverno.Palavras-chave: desfolhação, pastagem, produção de grãos Winter Cereals Submitted to Cuts in Double Purpose SystemABSTRACT -The winter cereals utilization in double purpose system allows to provide forage in critical food conditions periods and also to increase the income of production stablility by improving quality and produtivity of winter cereal grains. The experiment was conducted from April 1994 to September 1996, in Guarapuava, Paraná, to evaluate the potential for forage and grains of white oat (Avena sativa L.), wheat (Triticum aestivum), triticale (X. Triticosecale Witt.), black oat (Avena strigosa Schreb), rye (Secale cereale L.) and barley (Hordeum vulgare L.), seeking an use in conditions of double purpose. A randomized block experiment design, with treatments assigned to subdivided portions, in three replications, was used. In the plots the production systems (without cut, one and two cut) were studied and in the sub-plot, the genotypes. The two cut system was superior than the others concerning the dry matter, mainly the oat. For the grain yield, the systems without cut and one cut were superior, in spite of the greatest production of oat genotype under two cut. In all genotypes, the hectoliter weight increased and the mass of a thousand seeds decreased, when cuts were performed. Under apropriate handling conditions, considerable forage production can be reached, without affecting the posterior production of grains for winter cereals. IntroduçãoA região de Guarapuava, no Paraná, caracteriza-se pela alta aptidão à atividade agrícola. ...

This work aimed to evaluate the following hypotheses: 1) the daily grazing time (GT) and 2) forage searching are more associated with the sward structure than with the levels of daily forage allowance (FA). To this end we proposed a model that was tested through an analysis of the sward structure, grazing time, and displacement in grazing by heifers on the natural grassland of the Pampa Biome (southern Brazil), which has been managed by FA levels since 1986. For three seasons, between January 2009 and February 2010, we evaluated the effect of FA on the main descriptors of the sward structure (herbage mass, sward height, and tussocks frequency) and the effect of these on the GT, displacement rate (DR), and daily displacement (D) in grazing. The data were analyzed with the use of regression and descriptive analyses from three-dimensional contour graphs with the data of the sward structure and GT. The DR was not associated with the FA levels or sward structure; however, the DR presented a positive linear relationship with the D and GT. The incremental change in the GT was accompanied by an increase in the D. Lastly, independently of the level of the FA and season evaluated, the lower values of GT were always associated with the following structural configuration: forage mass between 1 400 and 2 200 kg DM Á ha À1 , sward height between 9 and 13 cm, and tussock levels not exceeding 35%. Outside these limits, a penalty occurred in the GT and displacement patterns of the heifers. We found evidence that a better understanding of the cause-effect relationships between the sward structure and the ingestive behavior of the animals demonstrates the possibility of increasing the performance of domestic herbivores with important economic and ecological consequences. Resumen El objetivo del estudio fue evaluar las siguientes hipó tesis: (i) si el tiempo de pastoreo diario (TP) y (ii) la búsqueda de forraje está n má s estrechamente relacionados a la estructura del pasto que a los niveles diarios de oferta de forraje (OF). Con este fin, propusimos un modelo que se puso a prueba en base al aná lisis de la estructura del pasto, el tiempo de pastoreo y el desplazamiento en pastoreo en terneras sobre un pastizal natural del Bioma Pampa (sur de Brasil) que, desde 1986, se ha manejado con distintos niveles de OF. En tresépocas, entre Ene/2009 y Feb/2010, se evaluó el efecto de la OF sobre los principales descriptores de la estructura del pasto (biomasa de forraje, altura y frecuencia de matas) y el efecto deéstos sobre el tiempo de pastoreo (TP), la tasa de desplazamiento (TD) y el desplazamiento diario (D). Los datos fueron analizados mediante regresió n y por aná lisis descriptivos a partir de grá ficos de contorno tridimensionales en base a los datos de estructura del pasto y TP. La TD no tuvo relació n con OF ni con la estructura del pasto, pero mostró una relació n lineal positiva con D. Incrementos en TP estuvieron asociados a incrementos en D. El estudio demostró la importancia de la estructura del pasto al constatar que, indep...

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Extensive livestock production in southern South America occupies ~0.5 M km2 in central-eastern Argentina, Uruguay and southern Brazil. These systems have been sustained for more than 300 years by year-long grazing of the highly biodiverse native Campos ecosystems that provides many valuable additional ecosystem services. However, their low productivity (~70 kg liveweight/ha per year), at least relative to values recorded in experiments and by best farmers, has been driving continued land use conversion towards agriculture and forestry. Therefore, there is a pressing need for usable, cost effective technological options based on scientific knowledge that increase profitability while supporting the conservation of native grasslands. In the early 2000s, existing knowledge was synthesized in a path of six sequential steps of increasing intensification. Even though higher productivity underlined that path, it was recognized that trade-offs would occur, with increases in productivity being concomitant to reductions in diversity, resilience to droughts, and a higher exposure to financial risks. Here, we put forward a proposal to shift the current paradigm away from a linear sequence and toward a flexible dashboard of intensification options to be implemented in defined modules within a farm whose aims are (i) to maintain native grasslands as the main feed source, and (ii) ameliorate its two major productive drawbacks: marked seasonality and relatively rapid loss of low nutritive value-hence the title “native grasslands at the core.” At its center, the proposal highlights a key role for optimal grazing management of native grasslands to increase productivity and resilience while maintaining low system wide costs and financial risk, but acknowledges that achieving the required spatio-temporal control of grazing intensity requires using (a portfolio of) complementary, synergistic intensification options. We sum up experimental evidence and case studies supporting the hypothesis that integrating intensification options increases both profitability and environmental sustainability of livestock production in Campos ecosystems.

Accurate estimates of bite mass and variations in the short-term intake rate of grazing herbivores has been historically considered as a fundamental methodological difficulty, a difficulty that increases with the complexity of the feeding environment. Improving these methodologies will help understand foraging behaviours in natural grazing conditions, where habitat structure and interactions among different forages influence feeding decisions and patterns. During the past 30 years, we have been developing the 'continuous bite-monitoring' method, an observational method that allows continuous assessment of foraging behaviours, including bite mass, instantaneous intake rate and food selection, in simple to complex feeding environments. The centrepiece of the method is a 'bite-coding grid' where bites are categorised by structural attributes of the forage to reflect differences in bite masses. Over the years, we have been using this method with goats, sheep, llamas and cattle across a range of different habitats. After reviewing the development of the method, we detail its planning and execution in the field. We illustrate the method with a study from southern Brazilian native Pampa grassland, showing how changes in the forages consumed by heifers strongly affect short-term intake rate during meals. Finally, we emphasise the importance of studying animals grazing in their natural environments to first identify the relevant processes that can later be tested in controlled experiments.