Highlights • Karelian spruces have morphology that is typical for P. obovata and characterized with genetic variation, described previously for P. abies. • Karelian spruces evolved due to introgression between P. abies and P. obovata. However, it is unclear whether Karelian spruces could be treated as P. fennica, because unequivocal morphological and genetic characters of the latter are still absent.
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