Palaeobiogeographical data on Cambrian trilobites obtained during the twentieth century are combined in this paper to evaluate palaeoceanographic links throughc.30 myr, once these arthropods biomineralized. Worldwide major tectonostratigraphic units are characterized at series intervals of Cambrian time and datasets of trilobite genera (629 for Cambrian Series 2, 965 for Cambrian Series 3, and 866 for the Furongian Series) are analysed using parsimony analysis of endemicity. Special attention is given to the biogeographical observations made in microcontinents and exotic terranes. The same is done for platform-basinal transects of well-known continental margins. The parsimony analysis of endemicity analysis resulted in distinct palaeogeographical area groupings among the tectonostratigraphic units. With these groupings, several palaeobiogeographical units are distinguished, which do not necessarily fit the previously proposed biogeographical realms and provinces. Their development and spatial distributions are broadly controlled by Cambrian palaeoclimates, palaeogeographical conditions (e.g. carbonate productivity and anoxic conditions) and ocean current circulation.Supplementary material:Global dataset of Cambrian Epoch 2 (A), Cambrian Epoch 3 (B) and the Furongian Epoch (C) trilobite genera are provided at:http://www.geolsoc.org.uk/SUP18669

The ptychoparioid trilobite Blainia gregaria Walcott, 1916 is described for the first time from the late middle Cambrian (Series 3) of the Argentine Precordillera. It occurs commonly in the upper part of La Laja Formation, a carbonate platform succession that crops out in the Precordillera Oriental of San Juan where it reaches a thickness of some 600 m. Sections were sampled on cerro Tres Marías in Sierra de Marquesado and in quebrada de Zonda at the northern termination of Sierra Chica de Zonda. More than 1200 specimens were recovered from 22 stratigraphic levels over a thickness of 155 m, located in the uppermost 15 m of the Soldano Member, the approximately 100 m thick Rivadavia Member, and in the lower 40 m of the Juan Pobre Member. Our biometric study dealt with 205 cranidia and 365 pygidia from cerro Tres Marías. Measured dimensions of a subset of this collection provide bivariate datasets that in some characters show broad scatter plots and relatively low linear correlation coefficients, indicative of rather wide intraspecific morphological variation. Six morphotypes (three cranidial and three pygidial), recognized mainly on the basis of shape, occur together in the same collections. This evidence for an unusually high degree of variability leads to a proposed synonymy of some 30 previously named species in North America belonging to two genera, Blainia Walcott, 1916 and Glyphaspis Poulsen, 1927, the latter considered a junior synonym. The biogeographic distribution of B. gregaria, as redefined here, shows that it was an endemic Laurentian species that inhabited the inner parts of the carbonate platform that rimmed the craton and mantled portions of its interior. Its presence in western Argentina is further evidence for the very close faunal relationship between the Precordilleran terrane, known as Cuyania, and Laurentia during the Cambrian.

The Lower Ordovician La Silla Formation of the Precordillera of west‐central Argentina is part of the west‐facing early Palaeozoic, tropical carbonate platform succession that comprises the core of the Cuyania terrane. Up to 360 m thick, it is exposed in several thrust sheets over a distance of some 250 km along and across depositional strike over a palinspastically unrestored distance of about 35 km. La Silla Formation is a strikingly pure limestone with subordinate finely crystalline dolomite and rare chert. It accumulated on a more or less uniformly subsiding passive margin. Copious precipitation of microcrystalline calcite, probably influenced by microbial activity to varying degrees, led to the generation of peloids, ooids and aggregates of these grains, as well as small amounts of lime mud, intraclasts, stromatolites and thrombolites. Rare bioclasts are limited mostly to scattered gastropods and trilobite sclerites; bioturbation is present locally. The array of carbonate rock types is grouped into eight recurring lithofacies, in order of decreasing abundance: (i) peloidal grainstone; (ii) laminated dolostone; (iii) intraclastic rudstone; (iv) microbial laminite; (v) peloidal packstone; (vi) ooidal grainstone; (vii) thrombolite boundstone; and (viii) mudstone. These facies represent sediments that formed solely in a shallow subtidal marine environment, with no evidence of restricted conditions, hypersalinity or subaerial exposure. No events of eustatic sea‐level change are recorded. By far the dominant facies is grainstone composed of well‐sorted, fine sand‐sized peloids and peloidal aggregates in homogeneous, tabular to gently undulating, medium to thick beds; cross‐lamination is scarce. Clusters of sub‐metre‐sized microbial patch reefs developed sporadically. The shallow platform is envisaged to have been covered by extensive peloidal sand flats and low‐relief banks, and little lime mud was generated. The setting was probably microtidal and may not have been affected by strong trade winds. It was washed by frequent, relatively gentle wave action but without experiencing powerful storms. In the middle member, anomalous lenses of intraclastic rudstone and laminated dolostone occur as graded beds overlying sharply downcut scoured surfaces up to 20 cm deep; these are interpreted to indicate a phase when accretion was punctuated occasionally by tsunamis generated from rift‐faulting seaward of the platform margin. The remarkably uniform peloidal grainstone composition over a broad area shows that, given the appropriate combination of climate, environmental and ecological factors, large portions of some early Palaeozoic platforms were dominated by grainy sediment and remained under well‐agitated conditions within fair‐weather wave‐base, without distinct lateral facies differentiation or tidal‐flat aggradation.

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Agnostoid trilobites from the Carlos Rusconi collections in the Museum of Natural Sciences of Mendoza, Argentina, are revised. The agnostoids were collected in the southern Precordillera of Mendoza Province, western Argentina. Revised taxa include Agnostus microcephalus (Rusconi), Lotagnostus (Lotagnostus) peladensis? (Rusconi), Lotagnostus (Lotagnostus) lasherensis (Rusconi), Homagnostus pehuenchensis Rusconi, Kormagnostus seclusus (Walcott), Kormagnostus? beltensis (Lochman in Lochman and Duncan), Hypagnostus parvifrons (Linnarsson), Ptychagnostus cuyanus (Rusconi), Ptychagnostus aculeatus (Angelin), Lejopyge laevigata (Dalman), Tomagnostella exsculpta (Angelin), Diplagnostus planicauda (Angelin), Oedorhachis typicalis Resser, Clavagnostus calensis Rusconi and Clavagnostus repandus (Westergård in Holm and Westergård). Some taxa of uncertain affinities are recognized and new specimens assignated to Lotagnostus and Pseudagnostinae are illustrated. Hypagnostus parvifrons, L. laevigata and C. repandus are reported for the first time in South America. Global correlation with previously described agnostoid faunas indicates that the outcrops range from middle Middle Cambrian to latest Late Cambrian in age.