Many boreal tree stands are neither clearly even-aged nor clearly uneven-aged. The stands may undergo a series of stages, during which an even-aged stand is transformed into two-storied mixed stand, and finally to multistoried or uneven-aged stand structure. The species composition often changes during the succession of stand stages. This study developed models for stand dynamics that can be used in different stand structures and species compositions. The model set consists of species-specific individual-tree diameter increment and survival models, and models for ingrowth. Separate models were developed for Scots pine, Norway spruce, and hardwood species. The models were used in a growth simulator, to give illustrative examples on species influences and stand dynamics. Methods to simulate residual variation around diameter increment and ingrowth models are also presented. The results suggest that mixed stands are more productive than one-species stands. Spruce in particular benefits from an admixture of other species. Mixed species improve diameter increment, decrease mortality, and increase ingrowth. Pine is a more beneficial admixture than birch. Simulations showed that uneven-aged management of spruce forests is sustainable and productive, and even-aged conifer stands growing on medium sites can be converted into uneven-aged mixed stands by a series of strong high thinnings.
The choice between even- and uneven-aged forest management is a topical issue as the negative impacts of clear-felling are being increasingly criticized and the profitability of even-aged management has been questioned. This study compared these management systems in spruce and pine stands in terms of timber, carbon, and bilberry benefits, all of which can be predicted with reasonable accuracy and quantified in terms of money. Management was optimized by maximizing the total net present value (NPV) of the three benefits in a steady-state situation. The currently recommended type of even-aged management was also included in the comparisons. Uneven-aged management was the best in terms of the total NPV and with respect to bilberry benefits (NPV of bilberry harvesting). It was also better than even-aged management in terms of timber benefits when the discount rate was more than 1%. The ranking was less clear in terms of carbon sequestration and discounted carbon benefits.
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