The skeletal record of tyrannosaurids is well-documented, whereas their footprint record is surprisingly sparse. There are only a few isolated footprints attributed to tyrannosaurids and, hitherto, no reported trackways. We report the world’s first trackways attributable to tyrannosaurids, and describe a new ichnotaxon attributable to tyrannosaurids. These trackways are from the Upper Cretaceous (Campanian - Maastrichtian) of northeastern British Columbia, Canada. One trackway consists of three tridactyl footprints, and two adjacent trackways consist of two footprints each. All three trackways show animals bearing southeast within an 8.5 meter-wide corridor. Similarities in depth and preservation of the tyrannosaurid tracks indicate that these three trackways were made by track-makers walking concurrently in the same direction. These trackways add significantly to previous osteology-based hypotheses of locomotion and behavior in Tyrannosauridae by providing ichnologic support for gregariousness in tyrannosaurids, and the first record of the walking gait of tyrannosaurids.

The collection and dissemination of vertebrate ichnological data is struggling to keep up with techniques that are becoming commonplace in the wider palaeontological field. A standard protocol is required to ensure that data is recorded, presented and archived in a manner that will be useful both to contemporary researchers, and to future generations. Primarily, our aim is to make the 3D capture of ichnological data standard practice, and to provide guidance on how such 3D data can be communicated effectively (both via the literature and other means) and archived openly and in perpetuity. We recommend capture of 3D data, and the presentation of said data in the form of photographs, false‐colour images, and interpretive drawings. Raw data (3D models of traces) should always be provided in a form usable by other researchers (i.e. in an open format). If adopted by the field as a whole, the result will be a more robust and uniform literature, supplemented by unparalleled availability of datasets for future workers.

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BackgroundFossil tracks made by non-avian theropod dinosaurs commonly reflect the habitual bipedal stance retained in living birds. Only rarely-captured behaviors, such as crouching, might create impressions made by the hands. Such tracks provide valuable information concerning the often poorly understood functional morphology of the early theropod forelimb.Methodology/Principal FindingsHere we describe a well-preserved theropod trackway in a Lower Jurassic (∼198 million-year-old) lacustrine beach sandstone in the Whitmore Point Member of the Moenave Formation in southwestern Utah. The trackway consists of prints of typical morphology, intermittent tail drags and, unusually, traces made by the animal resting on the substrate in a posture very similar to modern birds. The resting trace includes symmetrical pes impressions and well-defined impressions made by both hands, the tail, and the ischial callosity.Conclusions/SignificanceThe manus impressions corroborate that early theropods, like later birds, held their palms facing medially, in contrast to manus prints previously attributed to theropods that have forward-pointing digits. Both the symmetrical resting posture and the medially-facing palms therefore evolved by the Early Jurassic, much earlier in the theropod lineage than previously recognized, and may characterize all theropods.

Public engagement and the promotion of science to a wider non-academic audience form an integral role of the professional scientist in the twenty-first century. The high level of public interest in palaeontology means that the Earth's prehistoric past can provide an important medium through which to communicate information concerning contemporary scientific issues. Here we explain how modern computer techniques can be used to enhance public understanding of complex palaeontological issues.

Literature concerning dinosaur footprints or trackways exhibiting abnormal gait or morphology reflecting pathology (ichnopathology) is rare. We report on a number of Jurassic and Cretaceous occurrences of theropod footprints from western North America with unusual morphologies interpreted herein as examples of inferred pathologies, or ichnopathologies. The majority of ichnopathologies are primarily manifested in the digit impressions and include examples of swelling, extreme curvature, dislocation or fracture, and amputation. A number of occurrences are single tracks on ex situ blocks with substantial deformation (inferred dislocation or fracture), or absence of a single digit impression. Two occurrences are from in situ natural mould trackways, one of which is a lengthy trackway of a presumed allosauroid with no noticeable deformation of the digits or feet but with strong inward rotation of the left footprint toward the midline and a pronounced, waddling limp. The other is a tyrannosaurid trackway consisting of three footprints (one right, two left) with the two left prints exhibiting repetitive ichnopathology of a partially missing Digit II impression.Keywords Dinosaur tracks, Amputation, Antalgic gait, Compensated gait, Jurassic, Cretaceous, Ichnopathology, Laterality, Irregular gait, Limping gait, Shuffling gait, Waddling gait INTRODUCTIONOver the course of the past few decades, there has been a marked increase in the literature of pathology of dinosaurs (e.g., Tanke and Rothschild, 2002 and references therein). However, the literature on inferred pathology from dinosaur tracks and trackways (ichnopathology) has not seen a corresponding increase. Reports on ichnopathology in dinosaurs have been infrequent, with perhaps less than one paper published per decade since the beginning of modern studies of dinosaur traces in the early 19th century. It is possible that reports of ichnopathology have not kept pace with reports of pathologies on osteological specimens due to the various influences of track-bearing substrate, the behavior of the trackmaker, as well as a range of taphonomic factors that make such interpretations difficult. The sparse publication record of ichnopathology may also be a reflection of the cautious approach ichnologists employ when interpreting phenomena related to tracks.Even with a restrained approach toward footprint and trackway interpretation, some dinosaur tracks and trackways exhibit morphologies that have defied explanation by conventional behaviors of the trackmaker, or by the physical and dynamic qualities of the track surface, both pre-and post-burial.This article discusses previous reports of dinosaur ichnopathology with reinterpretations where such is warranted. Several recent and new reports of dinosaur ichnopathologies are described, from sites in the Jurassic of China and the Cretaceous of North America. A number of criteria are proposed that will aid in the interpretation of ichnopathologies

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