The collection and dissemination of vertebrate ichnological data is struggling to keep up with techniques that are becoming commonplace in the wider palaeontological field. A standard protocol is required to ensure that data is recorded, presented and archived in a manner that will be useful both to contemporary researchers, and to future generations. Primarily, our aim is to make the 3D capture of ichnological data standard practice, and to provide guidance on how such 3D data can be communicated effectively (both via the literature and other means) and archived openly and in perpetuity. We recommend capture of 3D data, and the presentation of said data in the form of photographs, false‐colour images, and interpretive drawings. Raw data (3D models of traces) should always be provided in a form usable by other researchers (i.e. in an open format). If adopted by the field as a whole, the result will be a more robust and uniform literature, supplemented by unparalleled availability of datasets for future workers.
Public engagement and the promotion of science to a wider non-academic audience form an integral role of the professional scientist in the twenty-first century. The high level of public interest in palaeontology means that the Earth's prehistoric past can provide an important medium through which to communicate information concerning contemporary scientific issues. Here we explain how modern computer techniques can be used to enhance public understanding of complex palaeontological issues.
Literature concerning dinosaur footprints or trackways exhibiting abnormal gait or morphology reflecting pathology (ichnopathology) is rare. We report on a number of Jurassic and Cretaceous occurrences of theropod footprints from western North America with unusual morphologies interpreted herein as examples of inferred pathologies, or ichnopathologies. The majority of ichnopathologies are primarily manifested in the digit impressions and include examples of swelling, extreme curvature, dislocation or fracture, and amputation. A number of occurrences are single tracks on ex situ blocks with substantial deformation (inferred dislocation or fracture), or absence of a single digit impression. Two occurrences are from in situ natural mould trackways, one of which is a lengthy trackway of a presumed allosauroid with no noticeable deformation of the digits or feet but with strong inward rotation of the left footprint toward the midline and a pronounced, waddling limp. The other is a tyrannosaurid trackway consisting of three footprints (one right, two left) with the two left prints exhibiting repetitive ichnopathology of a partially missing Digit II impression.Keywords Dinosaur tracks, Amputation, Antalgic gait, Compensated gait, Jurassic, Cretaceous, Ichnopathology, Laterality, Irregular gait, Limping gait, Shuffling gait, Waddling gait INTRODUCTIONOver the course of the past few decades, there has been a marked increase in the literature of pathology of dinosaurs (e.g., Tanke and Rothschild, 2002 and references therein). However, the literature on inferred pathology from dinosaur tracks and trackways (ichnopathology) has not seen a corresponding increase. Reports on ichnopathology in dinosaurs have been infrequent, with perhaps less than one paper published per decade since the beginning of modern studies of dinosaur traces in the early 19th century. It is possible that reports of ichnopathology have not kept pace with reports of pathologies on osteological specimens due to the various influences of track-bearing substrate, the behavior of the trackmaker, as well as a range of taphonomic factors that make such interpretations difficult. The sparse publication record of ichnopathology may also be a reflection of the cautious approach ichnologists employ when interpreting phenomena related to tracks.Even with a restrained approach toward footprint and trackway interpretation, some dinosaur tracks and trackways exhibit morphologies that have defied explanation by conventional behaviors of the trackmaker, or by the physical and dynamic qualities of the track surface, both pre-and post-burial.This article discusses previous reports of dinosaur ichnopathology with reinterpretations where such is warranted. Several recent and new reports of dinosaur ichnopathologies are described, from sites in the Jurassic of China and the Cretaceous of North America. A number of criteria are proposed that will aid in the interpretation of ichnopathologies
Relationships between non-avian theropod dinosaurs and extant and fossil birds are a major focus of current paleobiological research. Despite extensive phylogenetic and morphological support, behavioural evidence is mostly ambiguous and does not usually fossilize. Thus, inferences that dinosaurs, especially theropods displayed behaviour analogous to modern birds are intriguing but speculative. Here we present extensive and geographically widespread physical evidence of substrate scraping behavior by large theropods considered as compelling evidence of “display arenas” or leks, and consistent with “nest scrape display” behaviour among many extant ground-nesting birds. Large scrapes, up to 2 m in diameter, occur abundantly at several Cretaceous sites in Colorado. They constitute a previously unknown category of large dinosaurian trace fossil, inferred to fill gaps in our understanding of early phases in the breeding cycle of theropods. The trace makers were probably lekking species that were seasonally active at large display arena sites. Such scrapes indicate stereotypical avian behaviour hitherto unknown among Cretaceous theropods, and most likely associated with terrirorial activity in the breeding season. The scrapes most probably occur near nesting colonies, as yet unknown or no longer preserved in the immediate study areas. Thus, they provide clues to paleoenvironments where such nesting sites occurred.
The Upper Cretaceous Cantwell Formation in Denali National Park and Preserve (DENA), Alaska, contains an unparalleled fossil avian biodiversity. The Cantwell Formation, thousands of metres thick, was deposited near its current latitude and is exposed throughout much of DENA and elsewhere in the central Alaska Range. The Formation comprises a lower, dominantly fluvial sedimentary unit and an upper, mostly volcanic unit. Sedimentation of the lower unit was mainly in alluvial fan, braided and meandering stream, and lacustrine environments, with possible marginal-marine influence at times. Pollen data suggest that these sedimentary rocks are late Campanian or early Maastrichtian in age; thus the Cantwell Formation is correlative with other well-known dinosaur localities in Alaska. Bird tracks are preserved in multiple locations along a 40-km transect in DENA in fluvial and lacustrine deposits. Some bird tracks are found in association with dinosaur tracks and others are found on beds interbedded with dinosaur track-bearing layers. The approximate body sizes of the birds based on tracks show a range from sparrow-to heron-sized birds (∼25-30% larger than the modern Sandhill Crane Grus canadensis). The Cantwell Formation contains footprints assigned to several ichnotaxa found in either Asia or North America based on such morphological criteria as the presence or absence of a hallux, print size and shape, and angle of divarication: Aquatilavipes swiboldae, Ignotornis mcconnelli, Magnoavipes denaliensis sp. nov., Gruipeda vegrandiunus sp. nov. and Uhangrichnus chuni. The presence of a mixed Asian and North American ichnofauna suggests that at least some birds used Alaska as a bridge between Asia and North America. This diverse assemblage of avian traces, combined with the known fossil bone record and invertebrate trace fossil record, demonstrates that the northern Late Cretaceous polar region contained significant biodiversity.
The Wealden exposures on the Isle of Wight have long been noted for their dinosaur footprints and trackways and represent a unique ichnological resource. However, with a few notable exceptions, these ichnites remain largely unstudied and documentation has been sporadic and often concentrated on a few particular sites. Thus, their context within the Lower Cretaceous is poorly understood. The vertebrate ichnological record of the Wessex Sub-basin is currently being re-assessed. We review the main sites containing dinosaur footprints on the Isle of Wight. We also look at previously-known sites, and present new research using techniques such as photogrammetry to accurately record, preserve, and distribute ichnological data, especially data recorded in the dynamic foreshore and cliff environments, where many trace fossils are frequently lost as a result of human activity, weathering, erosion, and changing marine dynamics. Although the ichnoassemblages of the island's Wealden facies are dominated by ornithopod tracks including Caririchnium and Amblydactylus, we also report the first occurrence of the ankylosaurian track Tetrapodosaurus in both the Wessex and Vectis Formations, alongside the previously reported stegosaurian Deltapodus, as well as the occurrence of tracks left by saurichian tracemakers.
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