Despite being a fundamental aspect of biodiversity, little is known about what controls species range sizes. This is especially the case for hyperdiverse organisms such as plants. We use the largest botanical data set assembled to date to quantify geographical variation in range size for ∼ 85 000 plant species across the New World. We assess prominent hypothesised range-size controls, finding that plant range sizes are codetermined by habitat area and long- and short-term climate stability. Strong short- and long-term climate instability in large parts of North America, including past glaciations, are associated with broad-ranged species. In contrast, small habitat areas and a stable climate characterise areas with high concentrations of small-ranged species in the Andes, Central America and the Brazilian Atlantic Rainforest region. The joint roles of area and climate stability strengthen concerns over the potential effects of future climate change and habitat loss on biodiversity.

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Seasonally, snow‐covered forests are a critical source of water in the Western United States and are subject to major disturbances, including fire, harvest, disease and insect‐caused mortality, that have relatively unknown effects on water availability. In this study, we investigated changes in winter season snow accumulation and ablation in a forest following the Las Conchas fire in the Jemez Mountains of New Mexico. We investigated two competing sets of processes that should determine the peak annual snowpack prior to snowmelt: (1) reduced interception by forest canopy results in greater new snow accumulation and (2) increased winter season ablation of the snowpack results in reduced peak snowpack volumes. These processes were evaluated with approximately 800 spatially distributed manual observations of new snow, 1500 manual observations of peak snowpack, and light detection and ranging‐derived snow depth, vegetation and terrain datasets collected prior to the fire. A single snowfall event yielded significantly larger snow depths in the post‐burn area versus the unburned area (p < 0.001), with 25% to 45% interception in the unburned area and near zero in the post‐burn area. Conversely, the peak snowpack depths were significantly larger in the unburned area compared with the post‐burn area (mean of 55 and 47 cm, respectively), despite nearly identical peak snowpacks prior to the fire (72 and 72 cm, respectively). The lack of strong vegetation controls led to less variability at peak snowpack in the post‐burn area and a shift towards topographically controlled variability, caused by differences in elevation and aspect, occurring at larger spatial scales. The unburned area had roughly 10% more water available for melt than the post‐burn area, with winter season ablation reducing snowpacks by nearly 50% prior to melt in the post‐burn area. The relative importance of shortwave radiation to the snowpack energy balance and sublimation suggests that the 10% reductions in peak snow water storage found in these north‐facing areas could be a conservative estimate for winter season ablation following fire. Further work is necessary to assess the role that topography plays in altering water partitioning following forest disturbance and the potential implications for ecological health and downstream water resources. Copyright © 2013 John Wiley & Sons, Ltd.

The results indicate that wide-ranging species are indeed characterized by greater intraspecific variation and that species' phenotypes shift along environmental gradients in the same direction as the community phenotypes. However, across species, the rate of intraspecific trait change, reflecting plastic and/or adaptive changes across populations, is limited and prevents species from adjusting to environmental gradients as quickly as interspecific changes resulting from community assembly.