What determines our views on taxation and crime, healthcare and religion, welfare and gender roles? And why do opinions about these seemingly disparate aspects of our social lives coalesce the way they do? Research over the last 50 years has suggested that political attitudes and values around the globe are shaped by two ideological dimensions, often referred to as economic and social conservatism. However, it remains unclear why this ideological structure exists. Here, we highlight the striking concordance between these two dimensions of ideology and two key aspects of human sociality: cooperation and group conformity. Humans cooperate to a greater degree than our great ape relatives, paying personal costs to benefit others. Humans also conform to group-wide social norms and punish norm violators in interdependent, culturally marked groups. Together, these two shifts in sociality are posited to have driven the emergence of large-scale complex human societies. We argue that fitness trade-offs and behavioural plasticity have maintained strategic individual differences in both cooperation and group conformity, naturally giving rise to the two dimensions of political ideology. Supported by evidence from psychology, behavioural genetics, behavioural economics, and primatology, this evolutionary framework promises novel insight into the biological and cultural basis of political ideology.

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Political conservatives' opposition to COVID-19 restrictions is puzzling given the well-documented links between conservatism and conformity, threat sensitivity, and pathogen aversion. We propose a resolution based on the Dual Foundations Theory of ideology, which holds that ideology comprises two dimensions, one reflecting trade-offs between threat-driven conformity and individualism, and another reflecting trade-offs between empathy-driven cooperation and competition. We test predictions derived from this theory in a UK sample using individuals' responses to COVID-19 and widely-used measures of the two dimensions – ‘right-wing authoritarianism’ (RWA) and ‘social dominance orientation’ (SDO), respectively. Consistent with our predictions, we show that RWA, but not SDO, increased following the pandemic and that high-RWA conservatives do display more concerned, conformist, pro-lockdown attitudes, while high-SDO conservatives display less empathic, cooperative attitudes and are anti-lockdown. This helps explain paradoxical prior results and highlights how a focus on unidimensional ideology can mask divergent motives across the ideological landscape.

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This is a pre-print of a chapter that will appear in the forthcoming book, tentatively titled "A Research Agenda for Experimental Economics" edited by Ananish Chaudhuri, and has not yet been peer-reviewed.

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