Fifteen Recent species and three Miocene species of Rhysodini (Coleoptera: Carabidae: Scaritinae) are named and described as new species: Yamatosa phuka (Thailand), Arrowina nan (Thailand), Arrowina taksar (Nepal), Omoglymmius (Omoglymmius) truncatus (Thailand), Omoglymmius (Omoglymmius) batantae (Irian Jaya), Rhyzodiastes (Temoana) orestes (Tibet), Rhyzodiastes (Rhyzostrix) ininius (Guyane), Rhyzodiastes (Rhyzostrix) exsequiae (Brazil), Clinidium (Clinidium) ashei (Panama), Clinidium (Clinidium) brusteli (Ecuador), Clinidium (Clinidium) chandleri (Costa Rica), Clinidium (Clinidium) dux (Philippines), Clinidium (Clinidium) erwini (Costa Rica), Clinidium (Clinidium) felix (Panama), and Clinidium (Clinidium) talamanca (Costa Rica), and the Dominican Republic amber fossils, Clinidium (Clinidium) poinari, Plesioglymmius (Ameroglymmius) zayasi, and Clinidium (Tainoa) grimaldii. The males of Rhyzodiastes (Temoana) denticauda R. Bell and J. Bell and Clinidium (Clinidium) segne R. Bell and J. Bell, previously known only from females, are described. The females of Yamatosa schawalleri R. Bell and J. Bell and Clinidium (Clinidium) impressum R. Bell and J. Bell, previously known from males, are described. Range extensions or clarifications are given for the following 30 species: Leoglymmius lignarius (Olliff), Medisores abditus R. Bell and J. Bell, Dhysores thoreyi (Grouvelle), Rhysodes sulcatus Fabricius, Yamatosa boysi (Arrow), Yamatosa reitteri (Bell), Yamatosa sinensis R. Bell and J. Bell, Plesioglymmius (Juxtaglymmius) jugatus R. Bell and J. Bell, Omoglymmius (Orthoglymmius) cavifrons (Grouvelle), Omoglymmius (Orthoglymmius) crenatus (Grouvelle), Omoglymmius (Omoglymmius) laticeps R. Bell, Omoglymmius (Omoglymmius) germari Ganglbauer, Omoglymmius (Omoglymmius) vicinus (Grouvelle), Grouvellina tubericeps (Fairemaire), Grouvellina radama R. Bell and J. Bell, Rhyzodiastes (Temoana) preorbitalis R. Bell and J. Bell, Rhyzodiastes (Temoana) denticauda R. Bell and J. Bell, Clinidium (Mexiclinidium) championi R. Bell and J. Bell, Clinidium (Mexiclinidium) newtoni R. Bell and J. Bell, Clinidium (Clinidium) centrale Grouvelle, Clinidium (Clinidium) granatense Chevrolat, Clinidium (Clinidium) insigne Grouvelle, Clinidium (Clinidium) integrum Grouvelle, Clinidium (Clinidium) jolyi R. Bell and J. Bell, Clinidium (Clinidium) mathani Grouvelle, Clinidium (Clinidium) moldenkei R. Bell and J. Bell, Clinidium (Clinidium) oberthueri Grouvelle, Clinidium (Clinidium) rossi R. Bell, Clinidium (Clinidium) segne R. Bell and J. Bell, and Clinidium (Clinidium) spatulatum R. Bell and J. Bell. Four revised keys are presented: Key to Species of Genus Arrowina; Key to Genera of Subtribe Clinidiina; Key to Species of Subgenus Rhyzodiastes (Rhyzostrix); and Key to Species of genus Clinidium (Clinidium). A discussion of the phylogeny of Clinidium (Clinidium) is given. The terminology for the antennal setae of Omoglymmius americanus (Laporte), as used by Bell and Bell, is compared with that of Baker (2001).
Revue suisse Zool. Rhysodine Beetles in the Geneva collection: a new species of Yamatosa, and a major range extension for Omoglymmius sakuraii Nakane
The Cuban rhysodine fauna contains five species in two subtribes, two genera and three subgenera (possibly four, if Clinidium humeridens Chevrolat proves to belong to sensu stricto). Keys are given to all species including one member of subtribe Omoglymmiina, Plesioglymmius ( Ameroglymmius ) compactus Bell and Bell and four members of subtribe Clinidiina, Clinidium ( Protainoa ) extrarium Bell and Bell, Clinidium ( Tainoa ) curvicosta Chevrolat, and Clinidium ( Tainoa ) chevrolati Reitter. The subgenus of Clinidium humeridens Chevrolat is not known. The male of Plesioglymmius compactus Bell and Bell (synonymy, Rhysodes cuhanus Zayas), formerly unknown, is described for the first time. A specific locality for Clinidium extrarium Bell and Bell, previously of unknown country of origin, is also given here for the first time. ResumenLa entomofauna rhysodini de Cuba contiene cinco especies de dos subtribus, dos generos y tres subgeneros (tal vez cuatro, si Clinidium humeridens Chevrolat prueva pertenecer a sensu stricto). Llaves son descritas para todas las especies, incluyendo un miembro de la subtribu Omoglymmiina, Plesioglymmius ( Ameroglymmius ) compactus Bell y Bell y cuatros miembros de la subtribu Clinidiina, Clinidium (. Protainoa ) extrarium Bell y Bell, Clinidium ( Tainoa ) curvicosta Chevrolat y Clinidium ( Tainoa ) chevrolati Reitter. El subgenero de Clinidium humeridens Chevrolat no es conocido. El macho de Plesioglymmius compactus Bell y Bell (Sinonimia, Rhysodes cubanus Zayas) previamente desconocido es descrito por primera vez. Un lugar especifico de Clinidium extrarium Bell y Bell, previamente de pais desconocido, tambien es reportado aqui por primera vez.
The Rhysodini known from Australia are illustrated and described and keys are provided. Sloanoglymmiina, new subtribe, and Sloanoglymmius, new genus, are proposed for Rhysodes planatus Lea.The following new species are described (type localities indicated): Kaveinga (. Angekiva) stiletto (Queensland, Lamington National Park); Kaveinga (Angekiva) walfordi (Queensland, 29 km southeast of Mareeba); Rhyzodiastes (Rhyzoarca) ovicollis (Queensland, Ramsay's Scrub, Cooloola); Omoglymmius (Caeconavitia) okei (Queensland, Blue Mountains, Cape York Peninsula); Omoglymmius (5. str.) bituberculatus (Queensland, Boar Pocket); Omoglymmius (s. str.) monteithi (Queensland, West Claudie River, Iron Range). The Australian fauna contains 13 species in five subtribes, five genera and eight subgenera.Intraspecific variation, distributions within Australia, and zoogeographic relationships with other regions are given. Rhysodini are restricted to the forests along the east coast, including nearby moun¬ tains from Cape York to Tasmania. Sloanoglymmius is taxonomically isolated and shares only plesiomorphic characters with Leoglymmius. Both are restricted to Australia. Omoglymmius (Caeconavi¬ tia) previously thought to be endemic to Fiji, has a species in northern Queensland. Otherwise, the new taxa do not alter the picture of Australian rhysodine zoogeography as presented in previous papers. Annals of Carnegie Museum vol. 60Additional features include differentiation of a narrow, condyliform "neck", fu¬ sion and elongation of the mentum so that it conceals the remaining mouthparts in ventral view, and presence in males of "calcars", anteriorly directed processes on the apices of the hind, and usually also the middle tibiae. The calcars are absent in females, and thus are a reliable means of identifying the sexes. Since all known Rhysodini look red-brown in bright light and piceous to black if seen in dimmer light, color descriptions have been omitted. Rhysodini are limited to forested regions where there is sufficient rainfall to permit the decay of wood and the growth of slime molds. In general, they are absent from forests which have a strongly marked dry season. In Australia they seem to be restricted to wet or at least moderately wet sclerophyll and rain forest. They are difficult to find and collect. The most fruitful collection method involves the prying apart of suitable logs, stumps or roots with a screwdriver, small crowbar or similar instrument. At times, machetes, hatchets or saws are useful. Some species have been found in dead spots in recently fallen branches of live trees, in major roots as deep as three meters below the surface, or in rotten areas in the centers of large standing trees. Rhyzodiastes mirabilis is usually found on small sticks in forest litter (G. B. Monteith, personal communication, 1989).Rhysodini are often gregarious, although seemingly less so in Australia. If a single specimen is found within a log, a careful dissection of the log will often reveal a large number of specimens. It is sometimes practica...
Seven new species of Rhysodini (Coleoptera: Carabidae) are described, Kaveinga (Kaveinga) waai (Moluccas), Plesioglymmius {Juxtaglymmius) negara (Malay Peninsula), Omoglymmius {O.) emdomani (New Guinea), Rhyzodiastes (Temoana) riedeli (New Guinea), Rhyzodiastes (Temoana) mindoro (Philippines), Clinidium (Clinidium) onorei (Ecuador), Clinidium (C.) gilloglyi (Panama). Descriptions are given for either males or females of the following five species, previously known from only one sex; Omoglymmius (O.) pulvinatus ( Grouvelle), Clinidium (C.) howdenorum Bell and Bell, Clinidium (C.) dormans Bell and Bell, Clinidium (C.) crater Bell and Bell, Clinidium (C.) spatulatum Bell and Bell. An error in the description of Clinidium (C.) boroquense Bell is corrected. Range extensions or clarifications are given for the following 20 species: Arrowina anguliceps (Arrow), Yamatosa niponensis (Lewis), Yamatosa draco (Bell), Yamatosa sinensis Bell and Bell, Plesioglymmius {Ameroglymmius) reichardti, Omoglymmius (Orthoglymmius) coomani (Grouvelle), Omoglymmius (O.) sakuraii Nakane, Omoglymmius (O.) semperi Bell and Bell, Omoglymmius (O.) hiekei Bell and Bell, Omoglymmius {O.) bucculatus (Arrow), Omoglymmius (O.) patens Bell and Bell, Omoglymmius(0.) pulvinatus (Grouvelle), Omoglymmius (O.) sedlaceki Bell and Bell, Rhyzodiastes (Temoana) convergens Bell and Bell, Clinidium (Arctoclinidium) rosenbergi Bell, Clinidium (C.) insigne Grouvelle, Clinidium (C.) oberthueri Grouvelle, Clinidium (C.) rossi Bell, Clinidium (C.) moldenkei Bell and Bell, Clinidium (C.) sulcigaster Bell.
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