Rhysodine Beetles (Insecta: Coleoptera: Carabidae): New Species, New Data III

Abstract

Fifteen Recent species and three Miocene species of Rhysodini (Coleoptera: Carabidae: Scaritinae) are named and described as new species: Yamatosa phuka (Thailand), Arrowina nan (Thailand), Arrowina taksar (Nepal), Omoglymmius (Omoglymmius) truncatus (Thailand), Omoglymmius (Omoglymmius) batantae (Irian Jaya), Rhyzodiastes (Temoana) orestes (Tibet), Rhyzodiastes (Rhyzostrix) ininius (Guyane), Rhyzodiastes (Rhyzostrix) exsequiae (Brazil), Clinidium (Clinidium) ashei (Panama), Clinidium (Clinidium) brusteli (Ecuador), Clinidium (Clinidium) chandleri (Costa Rica), Clinidium (Clinidium) dux (Philippines), Clinidium (Clinidium) erwini (Costa Rica), Clinidium (Clinidium) felix (Panama), and Clinidium (Clinidium) talamanca (Costa Rica), and the Dominican Republic amber fossils, Clinidium (Clinidium) poinari, Plesioglymmius (Ameroglymmius) zayasi, and Clinidium (Tainoa) grimaldii. The males of Rhyzodiastes (Temoana) denticauda R. Bell and J. Bell and Clinidium (Clinidium) segne R. Bell and J. Bell, previously known only from females, are described. The females of Yamatosa schawalleri R. Bell and J. Bell and Clinidium (Clinidium) impressum R. Bell and J. Bell, previously known from males, are described. Range extensions or clarifications are given for the following 30 species: Leoglymmius lignarius (Olliff), Medisores abditus R. Bell and J. Bell, Dhysores thoreyi (Grouvelle), Rhysodes sulcatus Fabricius, Yamatosa boysi (Arrow), Yamatosa reitteri (Bell), Yamatosa sinensis R. Bell and J. Bell, Plesioglymmius (Juxtaglymmius) jugatus R. Bell and J. Bell, Omoglymmius (Orthoglymmius) cavifrons (Grouvelle), Omoglymmius (Orthoglymmius) crenatus (Grouvelle), Omoglymmius (Omoglymmius) laticeps R. Bell, Omoglymmius (Omoglymmius) germari Ganglbauer, Omoglymmius (Omoglymmius) vicinus (Grouvelle), Grouvellina tubericeps (Fairemaire), Grouvellina radama R. Bell and J. Bell, Rhyzodiastes (Temoana) preorbitalis R. Bell and J. Bell, Rhyzodiastes (Temoana) denticauda R. Bell and J. Bell, Clinidium (Mexiclinidium) championi R. Bell and J. Bell, Clinidium (Mexiclinidium) newtoni R. Bell and J. Bell, Clinidium (Clinidium) centrale Grouvelle, Clinidium (Clinidium) granatense Chevrolat, Clinidium (Clinidium) insigne Grouvelle, Clinidium (Clinidium) integrum Grouvelle, Clinidium (Clinidium) jolyi R. Bell and J. Bell, Clinidium (Clinidium) mathani Grouvelle, Clinidium (Clinidium) moldenkei R. Bell and J. Bell, Clinidium (Clinidium) oberthueri Grouvelle, Clinidium (Clinidium) rossi R. Bell, Clinidium (Clinidium) segne R. Bell and J. Bell, and Clinidium (Clinidium) spatulatum R. Bell and J. Bell. Four revised keys are presented: Key to Species of Genus Arrowina; Key to Genera of Subtribe Clinidiina; Key to Species of Subgenus Rhyzodiastes (Rhyzostrix); and Key to Species of genus Clinidium (Clinidium). A discussion of the phylogeny of Clinidium (Clinidium) is given. The terminology for the antennal setae of Omoglymmius americanus (Laporte), as used by Bell and Bell, is compared with that of Baker (2001).

Omoglymmius (s. str.) wukong sp. n., a new species from Xizang, China (Coleoptera, Rhysodidae, Omoglymmiini)

“…); taxonomic combination); Bell and Bell 1982: 207 (Omoglymmius ( sensu stricto ); redescription); Bell and Bell 1987: 685 (Omoglymmius ( s. str. ); distribution; remarks; key); Bell and Bell 2000: 74 (Omoglymmius ( s. str. ); distribution).…”

“…Bell and Bell (1982) excellently revised Omoglymmius and established eleven subgenera to classify the congeneric species. The nominotypical subgenus is the largest with 97 species (Lorenz 2005, Bell and Bell 2009, Hovorka 2015). However, in the fauna of East Asia, only two species in the subgenus Omoglymmius had been recorded before this study, namely O. ( s. str. )…”

Rhyzodiastes (Temoana) xii sp. nov. (Coleoptera: Rhysodidae: Clinidiini), <br />a new species from Hainan Island, China

“…Key to species of the Rhyzodiastes (Temoana) singularis species-group, modified based on Bell & Bell (1985, 2000 Habitus (Figs. 1A, B) elongate, rather narrow, and lustrous.…”

“…nov., belongs to the R. (T.) singularis species-group and was collected from Hainan Island, southeast China. An updated key to all species of the R. (T.) singularis species-group is compiled based on the version of Bell & Bell (1985, 2000 to include the new species.…”

‘Bellography’: Life and Contributions of Ross and Joyce Bell, two New England Naturalists

“…He spent the summer of 1956 collecting insects in Mexico with Don Van Horn, a graduate student companion at the University of Illinois, who had gone on to teach entomology at the University of Colorado. It was on this trip that Ross found his first undescribed rhysodine beetle species, and so began a thread of passionate discovery that established him as the world's expert on these interesting beetles (Bell 1970(Bell , 1973(Bell , 1975(Bell , 1977(Bell , 1979(Bell , 1985a(Bell ,b, 1998(Bell , 1999Bell and Bell 1962, 1975, 1978, 1979, 1981, 1982, 1987a,b,c, 1988, 1989b, 1991, 1993, 1995, 2000, 2002, 2009, 2010, 2011. In the following year Ross collected a new entomological companion through marriage to Joyce Elaine Rockenbach of Whitestone, Queens, New York City.…”