All Paleocene stages (i.e., Danian, Selandian and Thanetian) have formally ratified definitions, and so have the Ypresian and Lutetian Stages in the Eocene, and the Rupelian Stage in the Oligocene. The Bartonian, Priabonian and Chattian Stages are not yet formally defined. After the global catastrophe and biotic crisis at the CretaceousePaleogene boundary, stratigraphically important marine microfossils started new evolutionary trends, and on land the now flourishing mammals offer a potential for stratigraphic zonation. During the Paleogene the global climate, being warm until the late Eocene, shows a significant cooling trend culminating in a major cooling event in the beginning of the Oligocene, preparing the conditions for modern life and climate. Orbitally tuned cyclic sedimentation series, calibrated to the geomagnetic polarity and biostratigraphic scales, have considerably improved the resolution of the Paleogene time scale.
The GSSP for the base of the Eocene Series is located at 1. 58 m above the base of Section DBH in the Dababiya Quarry, on the east bank of the Nile River, about 35 km south of Luxor, Egypt. It is the base of Bed 1 of the Dababyia Quarry Beds of the El Mahmiya Member of the Esna Formation, interpreted as having recorded the basal inflection of the carbon isotope excursion (CIE), a prominent (3 to 5%) geochemical signature which is recorded in marine (deep and shallow) and terrestrial settings around the world. The Paleocene/Eocene boundary is thus truly a globally correlatable chronostratigraphic level. It may be correlated also on the basis of 1) the mass extinction of abyssal and bathyal benthic foraminifera (Stensioina beccariiformis microfauna), and reflected at shallower depths by a minor event; 2) the transient occurrence of the excursion taxa among the planktonic foraminifera (Acarinina africana, A. sibaiyaensis, Morozovella allisonensis); 3) the transient occurrence of the Rhomboaster spp. -Discoaster araneus (RD) assemblage; 4) an acme of the dinoflagellate Apectodinium complex. The GSSP-defined Paleocene/Eocene boundary is approximately 0.8 my older than the base of the standard Eocene Series as defined by the Ypresian Stage in epicontinental northwestern Europe.
A major gap in our knowledge of the evolution of marsupial mammals concerns the Paleogene of the northern continents, a critical time and place to link the early history of metatherians in Asia and North America with the more recent diversification in South America and Australia. We studied new exceptionally well-preserved partial skeletons of the Early Oligocene fossil Herpetotherium from the White River Formation in Wyoming, which allowed us to test the relationships of this taxon and examine its adaptations. Herpetotheriidae, with a fossil record extending from the Cretaceous to the Miocene, has traditionally been allied with opossums (Didelphidae) based on fragmentary material, mainly dentitions. Analysis of the new material reveals that several aspects of the cranial and postcranial anatomy, some of which suggests a terrestrial lifestyle, distinguish Herpetotherium from opossums. We found that Herpetotherium is the sister group to the crown group Marsupialia and is not a stem didelphid. Combination of the new palaeontological data with molecular divergence estimates, suggests the presence of a long undocumented gap in the fossil record of opossums extending some 45Myr from the Early Miocene to the Cretaceous.
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