We studied the effects of five thinning treatments (T1 = 5.5, T2 = 11, T3 = 16.5, T4 = 22.5 and T5 = 28.5 m(2) ha(-1) basal area under bark) x two fertilizer treatments (F0 = unfertilized and F1 = fertilized with 400 kg ha(-1) N plus 229 kg ha(-1) P) on growth and water relations of pole-sized Eucalyptus marginata J. Donn ex Sm. trees growing in southwestern Australia. Thinning reduced leaf area index (LAI) from 2.1 in the T4 and T5 treatments to 0.8 in the T1F0 treatment. Fertilizer had no effect on LAI in the T2, T4 or T5 treatments, but increased LAI by 45 and 20% in the T1 and T3 treatments, respectively. Thinning plus fertilizing increased diameter growth most in the fastest growing trees, from 0.4 cm year(-1) for trees in the T5F0 and T5F1 treatments to 0.7 and 1.2 cm year(-1) for trees in the T1F0 and T1F1 treatments, respectively. In both fertilizer treatments, stand basal area and volume growth increased with increasing stand density up to 15 m(2) ha(-1), and thereafter declined with increasing stand density, such that the growth rate of trees in the T5 treatment was only half of that at a stand density of 15 m(2) ha(-1). In response to fertilizer, growth rates of the slowest and fastest-growing trees increased from 0.35 and 3.5 m(2) ha(-1) year(-1) (F0) to 0.56 and 5.4 m(3) ha(-1) year(-1) (F1), respectively. Stand growth efficiency (growth per unit LAI) increased in response to thinning, and fertilizer increased stand growth efficiency at all stand densities. Throughout the dry season, T5 trees had lower predawn shoot water potentials (Psi(pd)) (minimum of -1.5 MPa) than T1 or T2 trees (minimum of -0.7 MPa). Fertilizer decreased Psi(pd) in T5 trees (by -0.9 and -1.5 MPa, respectively, in F0 and F1), but not in T1 or T2 trees. Stand growth rate was closely related to cumulative midday water stress (CMWS) over the dry season, and volume growth rate declined sharply from 6 m(3) ha(-1) year(-1) at a CMWS of 130 MPa days, to zero at a CMWS of 220 MPa days. Application of fertilizer to thinned stands increased LAI, stand growth efficiency and stand growth. In unthinned stands, fertilizer increased stand growth efficiency and stand growth; however, it also increased tree water stress, which limited the fertilizer-induced increases in LAI and growth. We attribute the increase in tree and stand growth in response to application of fertilizer to increased photosynthetic rates, increased allocation to stem wood, and in thinned stands also to higher LAIs.

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Chemical and mineralogical analyses of two representative laterite profiles developed from granite in south-westem Australia have shown that several depth zones defined by particular mineral suites and geochemistry may be recognized. These zones are not simply related to the conventional morphological pallid, mottled, and ferruginous zones.Changes in geochemistry in both profiles can be related to the stability of the major mineral species and to the common isomorphous substitutions occurring in them. Ca, Na, and Mg, which are mainly present in plagioclase feldspars and ferromagnesian minerals, are removed during the earliest stage of alteration of the parent granite. K, Mn, Zn, and residual Ca, Na, Mg, and Sr decrease as microcline and micas alter to kaolin. A1 and Fe are most abundant in the surface zones which contain secondary sesquioxide minerals.

The effects of an intense fire on the nutrient status of jarrah ^Eucalyptus marginata Donn ex Sm.) forest soils were investigated. At seven experimental sites, representing a range of soil and vegetation types, surface soils were sampled to a maximum depth of 20 cm before, immediately after, and 1 year after the fire.Immediately after the fire, concentrations of total and extracted nutrients in surface (0-3 cm) soils mcreased markedly. One year later, concentrations of extracted nutrients in surface soils had returned to levels close to pre-burn concentrations. During this period, some properties of sub-surface (3-20 cm) soils also changed: Bray-extractable phosphorus and electrolytic conductivity decreased, whereas e.xtractablepotassium andpH increased relative to pre-burn concentrations. Initial increases in concentrations of extractable forms of nutrients were determined largely by the amounts of nutrients deposited in fine ash, whereas the changes after 1 year depended more on the leaching of ions and their reactions with the mineral soil.Increases in total amounts of major nutrients in surface soils arising from additions offi ne ash were generally well correlated wtth amounts in pre-burn litter and with amounts in the components of organic matter likely to be consumed in the fire. Significant quantities of all nutrients are redistributed from aboveground sources to the soil surface during intense fires, including nutrients which are most susceptible to losses through volatilization. This redistribution of nutrients is discussed in relation to short-term and long-term effects on plant growth and functioning of the jarrah ecosystem.