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frame at t = 0, i.e. with its origin at (r0, ~00, z0) in our basic frame and with its x axis perpendicular to our basic z axis and pointing directly away from it. It can be shown that we should put It should be remembered that (rs, ~, z~) should refer to a left-handed helix in a right-handed frame.I should like to thank Mr G. Kreisel for a number of interesting discussions and in particular for suggesting the use of Parseval's theorem at a crucial point.

The DNA of higher organisms usually falls into two classes, one specific and the other comparatively nonspecific. It seems plausible that most of the latter originates by the spreading of sequences which had little or no effect on the phenotype. We examine this idea from the point of view of the natural selection of preferred replicators within the genome.

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