This study concerns the formation, taphonomy, and preservation of human footprints in microbial mats of present-day tidal-flat environments. Due to differences in water content and nature of the microbial mats and the underlying sediment, a wide range of footprint morphologies was produced by the same trackmaker. Most true tracks are subjected to modification due to taphonomic processes, leading to modified true tracks. In addition to formation of biolaminites, microbial mats play a major role in the preservation of footprints on tidal flats. A footprint may be consolidated by desiccation or lithification of the mat, or by ongoing growth of the mat. The latter process may lead to the formation of overtracks. Among consolidated or (partially) lithified footprints found on present-day tidal flats, poorly defined true tracks, modified true tracks, and overtracks were most frequently encountered while unmodified and well-defined true tracks are rather rare. We suggest that modified true tracks and overtracks make up an important percentage of fossil footprints and that they may be as common as undertracks. However, making unambiguous distinctions between poorly defined true tracks, modified true tracks, undertracks, and overtracks in the fossil record will remain a difficult task, which necessitates systematic excavation of footprints combined with careful analysis of the encasing sediment.
A new ichnospecies of a large theropod dinosaur, Megalosauripus transjuranicus, is described from the Reuchenette Formation (Early–Late Kimmeridgian, Late Jurassic) of NW Switzerland. It is based on very well-preserved and morphologically-distinct tracks (impressions) and several trackways, including different preservational types from different tracksites and horizons. All trackways were excavated along federal Highway A16 near Courtedoux (Canton Jura) and systematically documented in the field including orthophotos and laserscans. The best-preserved tracks were recovered and additional tracks were casted. Megalosauripus transjuranicus is characterized by tridactyl tracks with clear claw and digital pad impressions, and notably an exceptionally large and round first phalangeal pad on the fourth digit (PIV1) that is connected to digit IV and forms the round heel area. Due to this combination of features, M. transjuranicus clearly is of theropod (and not ornithopod) origin. M. transjuranicus is compared to other Megalosauripus tracks and similar ichnotaxa and other unassigned tracks from the Early Jurassic to Early Cretaceous. It is clearly different from other ichnogenera assigned to large theropods such as Eubrontes–Grallator from the Late Triassic and Early Jurassic or Megalosauripus–Megalosauropus–Bueckeburgichnus and Therangospodus tracks from the Late Jurassic and Early Cretaceous. A second tridactyl morphotype (called Morphotype II) is different from Megalosauripus transjuranicus in being subsymmetric, longer than wide (sometimes almost as wide as long), with blunt toe impressions and no evidence for discrete phalangeal pad and claw marks. Some Morphotype II tracks are found in trackways that are assigned to M. transjuranicus, to M.? transjuranicus or M. cf. transjuranicus indicating that some Morphotype II tracks are intra-trackway preservational variants of a morphological continuum of Megalosauripus transjuranicus. On the other hand, several up to 40 steps long trackways very consistently present Morphotype II features (notably blunt digits) and do not exhibit any of the features that are typical for Megalosauripus (notably phalangeal pads). Therefore, it is not very likely that these tracks are preservational variants of Megalosauripus transjuranicus or Megalosauripus isp. These trackways are interpreted to have been left by an ornithopod dinosaur. The high frequency of large theropod tracks in tidal-flat deposits of the Jura carbonate platform, associated on single ichnoassemblages with minute to medium-sized tridactyl and tiny to large sauropod tracks has important implications for the dinosaur community and for paleoenvironmental and paleogeographical reconstructions. As with most other known occurrences of Megalosauripus tracks, M. transjuranicus is found in coastal settings, which may reflect the preference of their theropod trackmakers for expanded carbonate flats where food was abundant.
Jurabrontes curtedulensis, a new ichnogenus and species of Late Jurassic giant theropod dinosaur track is described based on very well-preserved and morphologically-distinct tracks, all carefully excavated along federal highway A16 (Canton Jura, NW Switzerland). All trackways were systematically documented including parameter measurements, descriptions, outline drawings, orthophotos and laserscans. Jurabrontes is characterised by sub-equal track length and width, a small anterior triangle, weak mesaxony, three blunt digits (dII-III-IV) with pronounced (sub)triangular claw marks, a rounded heel, and clear phalangeal pad impressions. The combination of features of Jurabrontes is typical for a theropod (and not ornithopod) trackmaker. Jurabrontes is compared to other similar ichnotaxa and unnamed tracks of large theropods from the Early Jurassic to Late Cretaceous, from which it is clearly different. The sheer size of the largest tracks, that are amongst the largest worldwide and of similar size to Tyrannosauripus from the Late Cretaceous, suggests a 'megalosaurid' or large allosaurid theropod as a trackmaker. The presence of such large theropod tracks in tidal-flat deposits of the Jura carbonate platform and associated with small to large sauropod tracks has important palaeoecological implications for the dinosaur community and for palaeoenvironmental and palaeogeographical reconstructions.
The collection and dissemination of vertebrate ichnological data is struggling to keep up with techniques that are becoming commonplace in the wider palaeontological field. A standard protocol is required to ensure that data is recorded, presented and archived in a manner that will be useful both to contemporary researchers, and to future generations. Primarily, our aim is to make the 3D capture of ichnological data standard practice, and to provide guidance on how such 3D data can be communicated effectively (both via the literature and other means) and archived openly and in perpetuity. We recommend capture of 3D data, and the presentation of said data in the form of photographs, false‐colour images, and interpretive drawings. Raw data (3D models of traces) should always be provided in a form usable by other researchers (i.e. in an open format). If adopted by the field as a whole, the result will be a more robust and uniform literature, supplemented by unparalleled availability of datasets for future workers.
The Kimmeridgian shallow-water carbonates of the Swiss Jura Mountains display stratigraphic levels containing mass accumulations of nerineoidean gastropods. The macro-and microfacies of the rocks with these nerineoidean assemblages suggest that their occurrence was related to water depth, physical energy within the habitat, sedimentation rate, substrate stability, and food supply. Typical nerineoidean shells are high spired and exhibit unique spiral laminae (folds) on the internal walls. Mass accumulations consisting of large, heavy individuals are believed to represent a semisessile, suspensionfeeding community that utilized an external food source. During times of enhanced precipitation and run-off from the hinterland, terrestrial organic matter may have been supplied into the shallow sea, thus forming a source of food for the nerineoids. Alternatively, relative sea-level change during transgression may have led to largescale coastal erosion and flooding of the platform, also potentially leading to a mobilization of organic matter. Size and morphology of the high-spired shells seem to indicate a trend toward maximal external volume. Development of folds within the shell is interpreted as a modification to maintain a small internal volume. The faunal and floral composition of the strata indicates a suspension-rich, highenergy environment with a low-sedimentation rate and a stable substrate. This allowed mass colonization by epifaunal nerineoids, which, in turn, formed a frameworklike structure. The heavy shells acted as a grid, trapping the sediment in between. In this way, a reinforced carbonate accumulation formed.
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