Dinoflagellate cyst assemblages recovered from the La Meseta Formation cropping out in Seymour Island, Antarctic Peninsula, are studied herein and their distribution is compared with the biostratigraphic scheme available for the Palaeogene of the Southern Ocean and other high-latitude regions. In this way, the La Meseta Formation is dated as middle Lutetian to Priabonian (46.2–36 Ma), which differs from the age provided by other fossils, isotopes and also with the magnetostratigraphic scheme recently performed for the unit. The dinoflagellate cyst data support the proposal of ocean circulation patterns on the South American Shelf prior to the opening of Drake Passage. Assemblages from the La Meseta Formation contain Antarctic-endemic taxa which are also dominant in several circum-Antarctic sites, located south of 45° S. Their distribution reflects an ocean-circulation scheme with wide clockwise gyres surrounding Antarctica that were disrupted as a consequence of the deepening and definitive apertures of the Tasmanian Gateway and Drake Passage towards the Eocene/Oligocene transition. The palaeoenvironmental inference based on the S/D ratio (sporomorphs versus dinoflagellate cysts) and the P/G ratio (peridinioid versus gonyaulacoid dinoflagellate cysts) suggests an overall trend through the section from marine-dominated assemblages with poorly productive waters in the lower part of the section to more terrestrially dominated assemblages with increasing productivity in the upper part of the unit, reflecting a shallowing trend to the top.

The tectonic opening of the Tasmanian Gateway and Drake Passage represented crucial geographic requirements for the Cenozoic development of the Antarctic Circumpolar Current (ACC). Particularly the tectonic complexity of Drake Passage has hampered the exact dating of the opening and deepening phases, and the consequential onset of throughflow of the ACC. One of the obstacles is putting key regional tectonic events, recorded in southern Patagonian sediments, in absolute time. For that purpose, we have collected Campanian-Eocene sediment samples from the Chilean sector of Southern Patagonia. Using U-Pb radiometric dating on zircons and dinoflagellate cyst biostratigraphy, we updated age constraints for the sedimentary formations, and the hiatuses in between. Thick sedimentary packages of shallow-marine and continental sediments were deposited in the foreland basin during the early Campanian, mid-Paleocene, the Paleocene-Eocene boundary interval and the middle Eocene, which represent phases of increased foreland subsidence. We interpret regional sedimentary hiatuses spanning the late Campanian, early-to mid-Paleocene, mid-Eocene and latest Eocene-early Oligocene to indicate times of reduced foreland subsidence, relative to sediment supply. We relate these changes to varying subduction rates and Andean orogeny. Dinoflagellate cyst assemblages suggest that the region was under the influence of the Antarctic-derived waters through the western boundary current of the Subpolar Gyre, developed in the southwest Atlantic Ocean and thus argues for limited throughflow through the Drake Passage until at least the latest Eocene. However, the proliferation of dinoflagellate endemism we record in the southwest Atlantic is coeval with that in the southwest Pacific, and on a species level, dinoflagellate cyst assemblages are the same in these two regions. This suggests that both regions were oceanographically connected throughout the early Paleogene, likely through a shallow opening of a restricted Drake Passage. This implies a continuous surface-water connection between the south Pacific and the South Atlantic throughout the late Cretaceous-early Paleogene.

The study of microfloras and megafloras from western Argentina and southern Bolivia presented here extends our knowledge of the biodiversity and succession of floristic events during the Middle Devonian, and hence improves the current biostratigraphy. Among floral remains, species attributable to ‘Haplostigma’ are mostly recorded from the same Grandispora pseudoreticulata and other palynomorph-bearing sections at Balapuca (southern Bolivia) and Del Chaco and La Cortadera creeks in the Precordillera of Argentina. The northernmost record of Grandispora pseudoreticulata is recorded from a third palynoassemblage from the Pando x-1 corehole of northern Bolivia. A great similarity among all these assemblages (notably abundant Grandispora pseudoreticulata) suggests the identification of terrestrial connections, or at least proximity, of these local areas and other regions in South America during the Givetian. Comparison with other contemporary Gondwanan palynofloras shows cosmopolitan species (e.g. Geminospora lemurata, Samarisporites triangulatus, Archaeozonotriletes variabilis, Chelinospora concinna) along with some others with more restricted distribution (e.g. Grandispora pseudoreticulata, Leiotriletes balapucensis, Acinosporites ledundae). This pattern defines an Afro-South American Subrealm, which most likely results from the effects of palaeolatitude and, in a lesser way, local palaeoenvironmental conditions. On the other hand, such a level of cosmopolitanism supports previous palaeogeographical reconstructions where a narrow Rheic Ocean was developed between Euramerica and the northern parts of Africa and South America.

Micropaleontological and palynological samples from three Cenozoic dia− mictites at Cape Lamb, Vega Island, James Ross Basin were analysed. Fossiliferous sam− ples yielded reworked and autochthonous assemblages of Mesozoic calcareous nanno− fossils, impoverished Cretaceous foraminifera together with Neogene species, as well as Late Cretaceous dinoflagellate cysts, pollen, spores and abundant Cenozoic micro− foraminiferal linings. The recovered nannoflora indicates Early Cretaceous (Hauteri− vian-Albian) and Late Cretaceous (Santonian-Early Campanian) ages, suggesting an in− tensive reworking of marine sediments. The presence of the Early Cretaceous species Nannoconus circularis Deres et Acheriteguy in the diamictite represents its first record for the James Ross Basin. The scarce foraminiferal fauna includes Pullenia jarvisi Cushman, which indicates reworking from lower Maastrichtian-lower Paleocene sedi− ments, and also the Neogene autochthonous Trochammina sp. aff. T. intermedia. The in− ner−organic layer observed inside this specimen appears to be identical to microfora− miniferal linings recovered from the same sample. Palynomorphs found in the studied samples suggest erosion from the underlying Snow Hill Island and the López de Berto− dano Formation beds (upper Campanian-upper Maastrichtian). These recovered assem− blages indicate either different periods of deposition or reworking from diverse sources during Cenozoic glaciation, originating in James Ross Island and the Antarctic Peninsula with the influence of local sediment sources.

A proper understanding of the palaeoceanographic evolution of the Drake Passage during the Palaeogene is hampered by the lack of precise tools to date and correlate the sedimentary units of areas adjacent to the region. In this work, considering recently published radiometric U-Pb dates, we revised the age of a previous dinoflagellate zones for the middle to upper Eocene units of the Austral-Magallanes Basin. The quantitative analysis of middle to late Eocene dinoflagellate cyst assemblages from different localities close to the Drake Passage allowed us to reconstruct the palaeoenvironmental conditions and the possible surface ocean currents during this time in the area. Assemblages dated between 41.3 and 38.1 Ma represent relatively warm waters in inner shelf settings, while those ranged between 36 and 35 Ma reflect coastal areas with cool, nutrient-rich surface waters. The proposed surface ocean circulation pattern, based on dinoflagellate cysts distribution between 41.3 and 38.1 Ma, agrees with the results of a palaeoclimatic numerical model simulation performed with a Drake Passage shallow opening of 100 m depth. At c. 36 Ma, several Antarctic gonyaulacacean taxa tolerant to relatively warmer waters were replaced by some Antarctic peridinacean species better adapted to colder conditions. This change could be linked to a progressive deepening of the Drake Passage that is estimated to have reached 1000 m depth promoting a cooling in the South Atlantic. Such passage depth would have enabled stronger flows from the Pacific to the Atlantic Ocean, which is reflected by the increase of cosmopolitan species.

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Rich palynological assemblages have been obtained from the Middle Devonian Chigua Formation (Chinguillos Group, San Juan Province), western Precordillera, Argentina, at two new localities (Del Chaco and Don Agustín creeks). From the palyniferous levels at Del Chaco Creek, there are two assemblages: one is very rich in microplankton (assemblage 1) whereas the other is dominated by spores (assemblage 2). The level obtained from Don Agustín Creek, poorest in overall taxa, is also rich in microplankton (assemblage 3). Compared with coeval microfloras elsewhere, a late Emsian–early Eifelian, and an early Givetian age are proposed for the first and the second assemblages respectively. A third assemblage (assemblage 3) tentatively represents a timespan around the Givetian–Frasnian boundary. The proposed age based on the palynomorphs reinforces previous palaeontological records. Variation in the microplankton diversity, fluctuations in the microplankton/spore ratio and differences in acritarch morphology are used here to interpret changing proximity to palaeoshoreline and so to recognize fluctuations in relative sea level that occurred during the deposition of the Chigua Formation.

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