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Two versions of a stage-structured model of Cirsium vulgare population dynamics were developed. Both incorporated density dependence at one stage in the life cycle of the plant. In version 1 density dependence was assumed to operate during germination whilst in version 2 it was included at the seedling stage. Density-dependent parameter values for the model were estimated from annual census data in a factorial grazing experiment. Version 1 of the model produced significant estimates of density dependence under field conditions. The estimated values, when included in a simulation of the dynamics, produced two-point limit cycles under conditions of hard grazing. The limit cycles were most pronounced at the early rosette stage. Comparison of the effects of density dependence at the two different stages in the life cycle revealed a strong difference in predicted dynamics. This emphasizes the importance of determining where density dependence operates under field conditions and the potential problems of arbitrarily assigning it to particular life-history stages. Version 1 of the model produced a good prediction of observed mean plant density across the different grazing treatments (r =0.81, P<0.001).

The use of destocking as a means of promoting pasture recovery was evaluated on two important rangeland types of the semi-arid tropics of north west Australia. Recovery on eroded calcareous red soils was substantial within five years. Standing biomass and species composition, particularly the contribution of the major forage grass Enneapogon spp., was then similar to areas in good condition. However, an exotic shrub, Calotropls procera, invaded the area during the study. Grazing limited its increase to 200 bushes ha-1 compared to unstocked areas where it increased to 1,000 bushes ha-1. Destocking had no effect on cracking clay soils in good condition, dominated by Chrysopogon fallax and Iseilema fragile. An index based on similarity measures of herbage standing biomass, followed closely the course of recovery of the eroded calcareous soil during the period. There is a need for range condition assessment and the development and application of appropriate pasture management strategies for the whole district.