Begomoviruses have either a monopartite (single DNA) or a bipartite (two DNA components: DNA-A and DNA-B) genome organization, infecting mostly dicotyledonous plants. The DNA-A of bipartite and the single component of monopartite begomoviruses contain five (sometimes six) Open Reading Frames (ORFs), one (AV1) or two (AV1 and AV2) in the viral sense (V-sense) strand and four (AC1 to AC4) in the complementary sense (C-sense) strand. Both the DNA-A and
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