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Carbohydrates represent a low-cost energy source compared to others, thus being critical for reducing the costs associated with producing formulated diets (Gao et al., 2010; Mohanta, Mohanty, & Jena, 2007). In addition, it is interesting to include non-protein energy sources such as carbohydrates in diets so that fish can direct most of the dietary protein to growth, improving animal performance (Bautista, 1986; Gao et al., 2010). It is important to emphasize that starch granules are essential for the extrusion of rations, because they expand when subjected to heat, pressure and moisture, which allows fluctuation in the diets (Laif & Kokini, 1991). Corn bran, generally used in animal feed, is considered to be one of the most palatable ingredients among cereals, being mainly made up of starch (Pastore, Gaiotto, Ribeiro, & Nunes, 2013). In general, herbivorous and omnivorous fish are more able to use carbohydrates