Graphical Abstract Research HighlightsWe identified 8 heat shock proteins (3 small Hsps, 4 Hsp70's, Hsp90). Hsp70a (cytosolic clade) and Hsp21, and Hsp22 were induced by handling stress. Hsp70a expression was elevated in shallow, active animals. Hsp22 expression was elevated in deep, diapausing animals. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 *Research highlights

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