Digital technologies increasingly mediate relations between humans and nonhumans in a range of contexts including environmental governance, surveillance, and entertainment. Combining approaches from more-than-human and digital geographies, we proffer ‘digital ecologies’ as an analytical framework for examining digitally-mediated human–nonhuman entanglement. We identify entanglement as a compelling basis from which to articulate and critique digitally-mediated relations in diverse situated contexts. Three questions guide this approach: What digital technologies and infrastructures give rise to digital entanglement, and with what material consequences? What is at stake socially, politically, and economically when encounters with nonhumans are digitised? And how are digital technologies enrolled in programmes of environmental governance? We develop our digital ecologies framework across three core conceptual themes of wider interest to environmental geographers: (i) materialities, considering the infrastructures which enable digitally-mediated more-than-human connections and their socioenvironmental impacts; (ii) encounters, examining the political economic consequences and convivial potentials of digitising contact zones; (iii) governance, questioning how digital technologies produce novel forms of more-than-human governance. We affirm that digital mediations of more-than-human worlds can potentially cultivate environmentally progressive communities, convivial human–nonhuman encounters, and just forms of environmental governance, and as such note the urgency of these conversations.