“…In addition, recent studies of ancient human genomes showed continuous gene flow between Copper Age steppes and Caucasus peoples ( 3 , 4 , 76 , 77 ), and later, during the Bronze Age, between Mesopotamia, Anatolia, the southern and northern Caucasus, and the steppes ( 78 ). This exchange between human groups appears to intensify during a century-long period of cooling and desertification, known as the 4.2-ka (thousand year) event [e.g., ( 79 ) but see ( 80 )], which may have affected subsistence strategies and social networks in the steppe zone [e.g., ( 78 , 81 )]. On present evidence, this climatic event seems to be broadly contemporaneous with the arrival of nonlocal horse mitochondrial haplogroups and coat colors in the Caucasus and Anatolia and linked to the expansion of horse husbandry and possibly Indo-European languages [e.g., ( 14 , 75 ) but see ( 82 )].…”