“…Keenan et al (1998) revised the genus Scylla, combining genetic and morphometric data. Allozyme electrophoresis and analysis of mitochondrial DNA sequences (COI I and 16 S RNA) revealed four distinct species of Scylla, which are now generally accepted: S. serrata (Forskål, 1775), S. tranquebarica (Fabricius, 1789), S. olivacea (Herbst, 1796), and S. paramamosain (Estampador, 1949) (Fuseya & Watanabe, 1996;Keenan et al, 1998;Klinbunga et al, 2000;Imai et al, 2004;Fuseya et al, 2007;Lin et al, 2007;Ogawa et al, 2011;Mandal et al, 2014).…”