Drawing on a text-based ethnography of digital writing in a biology laboratory, this article examines the text trajectory of a scientific manuscript and a scientific team's related writing for public audiences, including for citizen scientists. Using data drawn from texts, observations, interviews, and related artifacts, the author examines how scientists conceptualize and adapt their multimodal writing for specialized scientific audiences as well as lay audiences interested in the work of scientific inquiry. Three conceptsmeaning compression, meaning expansion, and meaning attention-were used to analyze the multimodal strategies that scientists employ when composing for different audiences. Findings suggest that while scientists often restrict their writing practices to meaning compression to maintain the values and conventions of scientific genres, they also sometimes deploy a wider range of multimodal strategies when writing for nonspecialist audiences. These findings underscore the complex rhetorical environments scientists navigate and the need to support emerging scientific writers' development as versatile