“…If virus and grain surface charges are known, virus attachment behavior can be predicted, at least qualitatively, using classic DLVO theory (Derjaguin and Landau, 1941;Verwey and Overbeek, 1948). Conditions that reduce the electrostatic repulsion between viruses and grains, like low pH, presence of divalent cations, and high ionic strength, favor attachment (Zerda et al, 1985;Penrod et al, 1996).…”