Prior to domestication, livestock ancestors had coat colors that provided camouflage against potential predators. Wild species mostly had a uniform phenotype with species-specific coat colors and patterns (Charon & Lipka, 2015;Neves et al., 2017). With domestication, a large variation of phenotypes arose driven by adaption to different habitats and climate conditions as well as by different human preferences (Bellone, 2010;Cieslak et al., 2011;Neves et al., 2017).Numerous genes influencing pigmentation and coat color-associated loci have been identified (Cieslak et al., 2011). These genes often either influence production of the pigment melanin or its distribution by affecting the underlying pathways (Bellone, 2010;Hubbard et al., 2010). Two different types of melanin-eumelanin and phaeomelanin-are synthesized and stored in