“…Mismatches between stem water and available water sources might reflect spatial heterogeneity of water sources in the soil (Brunel, Walker, & Kennett-Smith, 1995) as well as limitations in our current understanding of soil water isotopic composition . Instead of a single well-mixed reservoir of soil moisture (Hewlett & Hibbert, 1967), soil water may be ecohydrologically partitioned into multiple, unmixed water pools that vary in age and mobility (McCutcheon, McNamara, Kohn, & Evans, 2017;Sprenger et al, 2019;Sprenger, Leistert, Gimbel, & Weiler, 2016). This incomplete mixing and compartmentalisation of water sources has been described as the "two-water worlds" hypothesis based on observations of plants that appear to utilize soil water with low mobility instead of water that follows preferential flow pathways (Brooks et al, 2010;McDonnell, 2014).…”