“…Lastly, evidence from in situ forests suggests that late Pennsylvanian forests, while taxonomically and vertically heterogeneous, had rather open canopy strata with spatially separated arborescent taxa resulting in little crown overlap (reviewed in DiMichele and Falcon-Lang, 2011, but see Opluštil et al, 2009Opluštil et al, , 2014. In contrast, Cisularian forests show evidence of denser communities, suggestive of more continuous canopy strata (Gulbranson et al, 2012;Wang et al, 2012;Luthardt et al, 2016), although less dense forests also occurred (Gulbranson et al, 2012;Opluštil et al, 2021). It could thus be argued that selective pressure for an aerial locomotion in a continuous canopy was minimal prior to the Cisularian, but this assumption requires further paleoecological studies, especially with respect to canopy density and continuity.…”