“…Candidate gene-based association analysis revealed that major genes controlling the flowering time, FLOWERING LOCUS C (FLC), FRIGIDA (FRI), VERNALIZATION-INSENSITIVE 3 (VIN3) and CRYPTOCHROME 2 (CRY2), are associated with natural variation (Shindo et al, 2005;Balasubramanian et al, 2006;Wollenberg and Amasino, 2012). After developing a number of accessions with the SNP information, the genome-wide association of genes regulating flowering time was identified by GWAS (Atwell et al, 2010;Li et al, 2014;Sasaki et al, 2015;The 1001Genomes Consortium, 2016Zan and Carlborg, 2018). GWAS on flowering time have been conducted in various crops other than Arabidopsis, leading to the discovery of new loci associated with flowering time (maize: Bouchet et al, 2013, rice: Huang et al, 2012, barley: Muñoz-Amatriaín et al, 2014, soybean: Zhang et al, 2015b, rapeseed: Xu et al, 2015, common bean: Moghaddam et al, 2016.…”