“…Starch can be modified by α glucan (Zhou et al ., 2017). When carbon shortage occurs, there are potentially multiple pathways by which starch may be degraded to soluble sugars, for example by α amylases, β amylases and other enzymes (Witt & Sauter, 1994; Smith, 2007; Zeeman et al ., 2010). Starch pathway studies in sink tissues have shown expression of enzymes involved in starch synthesis, as sucrose synthase (SUS), AGP and branching enzymes, and in starch degradation, such as starch phosphorylase, disproportionating enzymes (DPE), de‐branching enzymes, α and β amylases, maltose translocators and glucose translocators (Smith et al ., 2005; Geisler‐lee et al ., 2006; Regier et al ., 2009; Silva et al ., 2017; Noronha et al ., 2018).…”