“…Streamflow during low-flow periods largely originates from groundwater (e.g., Costelloe, Peterson, Halbert, Western, & McDonnell, 2015;Soulsby et al, 2007), including bedrock groundwater (e.g., Oda, Suzuki, Egusa, & Uchiyama, 2013;Sayama, McDonnell, Dhakal, & Sullivan, 2011;Tague & Grant, 2004). The properties of these groundwater stores, and thus their responses to extended dry periods, vary markedly from place to place, so that the magnitude of low flows can also be highly variable in space (e.g., Floriancic et al, 2018;Smakhtin, 2001;Sun, Kasahara, Otsuki, Saito, & Onda, 2017). Previous studies have shown that landscape characteristics, like topography (Kroll, Luz, Allen, & Vogel, 2004;Kuentz, Arheimer, Hundecha, & Wagener, 2017;Li et al, 2018;Sayama et al, 2011;Staudinger, Weiler, & Seibert, 2015), geology (Tague & Grant, 2004), land use (Ahn & Merwade, 2017;Smakhtin, 2001), or the extent of quaternary deposits and unconsolidated layers (Roy & Hayashi, 2009;Winkler et al, 2016) can affect the magnitude of low flows.…”