“…An alternative possibility is that savanna-forest boundaries may be highly dynamic over longer time-scales (Schwartz et al, 1986, West et al ., 2000, Staver et al , 2017, Aleman et al 2019) and in response to changing climate or climate events (Hirota el al , 2010, Breman et al, 2012, Beckett & Bond, 2019, Wright et al, 2021, Sato et al , 2021). Spatially explicit modelling work has argued that fires at the savanna-forest boundary can have major effects on the biogeographic distribution of biomes (Staver et al , 2011b, van de Leemput et al , 2015, Goel et al , 2020a, Goel et al , 2020b, Wuyts et al , 2019) and local observations confirm that boundaries are sometimes dynamic through time (Brook & Bowman, 2006, Silva et al , 2008, Ibanez et al , 2013, Beckett & Bond, 2019).…”